FRUIT OF MELOCANNA BAMBUSOIDES. 415 
For the epidermis (Pl. 47. fig. 55). besides having stomata, is differentiated into long 
and short cells (* Kurzzellen," Pfitzer) and two kinds of hairs, viz., fine, stiff- pointed, 
l-celled bristles, and smaller, thin-walled, 2-3-celled, obtuse, club-shaped hairs, which 
are usually bent above the basal cell so as to lie more or less on the epidermis, corre- 
sponding to the “Winkelhaare” of Grobb (* Anat. d. Epiderm. Gramineenblitt.,” in Bibl. 
Bot. vii. p. 20). As already mentioned, the coleoptile always bears a small bud ( Pl. 45. 
fig. 23; Pl. 47. figs. 46-48, 5.) with several distinct leaf-primordia in its axil, either 
opposite the radicle or more frequently slightly lateral. "The first leaf after the coleoptile 
(fig. 46, 1.) is exactly opposite to it, flattened and slightly 2-keeled on the back, not 
unlike an ordinary dorsal bract (** adossirtes Vorblatt,” Eichler), a condition evidently due 
to the pressure of the radicle, and disappearing as soon as the leaf grows beyond it in 
the course of germination. Like the coleoptile, the first leaf possesses numerous vascular 
bundles. The second leaf is so placed that its median plane cuts that of the coleoptile 
and first leaf at an angle of 30°; the third Jeaf is approximately opposite the second, 
The fourth diverges from the second by about 60°, and its median plane is therefore at a 
right angle to that of the coleoptile. The following leaves diverge approximately at an 
angle of 180° from the preceding leaf. The median plane of the coleoptile and the first 
leaf coincide therefore with that of the embryo, whilst the fourth and the following 
leaves of the plumule are placed at right angles to it, and the second and third leaves 
are in an intermediate position. I may add that the second leaf overlaps the third 
very distinctly on the outer side, whilst it is, though slightly, overlapped by it on the 
inner side (¢. e., the side towards the radicle). In the following leaves the overlapping 
is quite normal. 
2. The Radicle (Pl. 45. fig. 3, 7.; Pl. 47. figs. 36-43, rad.).—Like the plumule, the 
radicle has the shape of an oblique cone, but it is rather smaller. I have already 
mentioned that it consists, as is generally the case in Graminew, of a coleorrhiza, which 
is very large, a primary root (Pl. 47. fig. 46, r. I.), and a few secondary roots (Pl. 47, 
figs. 47, 51), which are arranged almost concentrically, and start at a level whioh 
corresponds to a cross-section from the inner angle of the embryo to the junction of 
scutellum and coleorrhiza on the opposite side. There is only one point in the structure 
of the radicle to which I wish to draw attention. As stated above, there is no epiblast 
in Melocanna, but on the outer side (i. e., the side opposite to that where we would 
-look for the epiblast) I once noticed a small scale (Pl. 47. figs. 39 ex., 40) of the shape 
and structure of an epiblast, whilst in another case only an irregular swelling was 
perceptible. In a third seed I found a short obtuse protuberance placed somewhat 
laterally and projecting downwards. At first I thought it might correspond to some 
secondary root, but, in dissecting it, I found it to consist of the same parenehy MA as 
the coleorrhiza, and that it was not correlated to any secondary root which was then 
discernible. Another peculiarity of the coleorrbiza is the pee of vere: d though 
comparatively deep furrows (Pl. 47. figs. 39, 53), extending longitudinally on : e upper 
and outer side. "There is apparently no definite rule as to their number an position, 
and, so far as Iwas able to see, no distinct correlation to anything in the internal 
