36 NORTH AMERICAN FLORA [VorumE 16 
narrowly marginate, rusty-puberulous, soon glabrescent; primary branches very short, 
knob-like, the terminal bud relatively large, usually abortive, closely invested with rusty 
hairs; sterile secondary (geminate) pinnae 12-30 cm. long, deltoid-ovate, inequilateral, 
bipinnate or at the base tripinnate, the rachis divaricate-flexuous; tertiary segments 2-4 
pairs, 2-5.5 cm. apart on each side, petiolate (1-2.3 cm.), gradually smaller toward the 
acuminate apex, the terminal segment lanceolate or narrowly subhastate from an unequally 
cuneate or obliquely lobate base, usually 3-5 cm. long, 1 cm. or less broad at the base, the 
middle segments mostly trifoliolate, the divisions similar to the terminal segment; lower- 
most tertiary segments up to 13 cm. long, rhombic-deltoid, with about 3 pairs of very dis- 
tant quaternary segments, these similar to the terminal tertiary segment, the lowermost 
often with a smaller conform free segment, distinctly stalked (5 mm.), the rachis strongly 
flexuous, delicately foliaceo-marginate; costules elevated, flexuous, extending to the apex ; 
veins very oblique, 1-2 times dichotomous, or in the single basal lobes subpinnate-flabel- 
late ; leaf-tissue membrano-herbaceous, bright clear green above, paler below, essentially 
glabrous, the margins everywhere marked by shallow crenations, these serrulate-dentate. 
Fertile secondary (geminate) pinnae similar in size, at least bipinnate, always tripin- 
nate at the base, the quaternary segments elongate-deltoid with the basal divisions short, 
narrowly cuneate, sessile or short-stalked, and often cleft nearly to the base into 2 unequal 
divisions; ultimate segments all deeply and very obliquely incised toward the base, above 
strongly serrate; sporangiophores usually single and terminal upon the lobes, oblong, up 
to 5 mm. long, obscurely pilose upon both surfaces; spores lightly tuberculate, the surfaces 
smooth. 
TYPE LOCALITY: Near Lake Miragoan, Haiti. 
DISTRIBUTION : Known only from the island of Santo Domingo. 
ILLUSTRATION: Plumier, Traité Foug. f/. 92. 
2. ACTINOSTACHYS Wall.; Hook. Gen. Fil. A/. 277, A. 1842. 
Fronds simple, linear, mostly triquetrous or flattish from a dark terete or semi-terete 
base, prominently unicostate. Sporangiophores terminal, the divisions elongate, spuriously 
digitate in a penicillate tuft from a very short inconspicuous prolongation of the costa. 
Sporangia in 2 rows, one upon each side of the costa, but apparently in 4 more or less com- 
plete rows by the bending of the crowded sori alternately to left and right; indusium 
continuous, formed of the narrowly reflexed margin of the segments. 
Type species, Acrostichum digitatum L. 
Fronds 5-20 em. long; fertile segments 3-8, 8-20 mm. long. 1. A. Germant. 
Fronds 25-55 cm. long; fertile segments 6-14, 2-4.5 cm. long. 2. A. Pennula. 
1. Actinostachys Germani Fée, Mém. Foug. 11: 123. 1866. 
Schizaea Germani Prantl, Schiz. 132. 1881. 
Plants from roundish or oblong-ovoid bristly tubers 1 cm. or less long (these sunk 9-12 
cm. in a substratum of humus or decayed wood), sessile or distant 1-3 cm. upon a slender 
ascending tortuous offshoot ; fronds one or several, very slender, 5-20 cm. long, terete or 
angled at the dark castaneous base, above strongly costate on the under surface, thus 
triquetrous, in the upper part flattish, about 1 mm. broad, glabrous; fertile segments 3-8, 
closely penicillate, subequal, 8-20 mm. long, slender, acuminate, the costa conspicuously 
pilose below ; sporangia in 2 rows, usually appearing to bein 4; spores delicately maculate. 
TyPE LOCALITY: Sainte-Rose, Petit-Bourg, Guadeloupe. 
DISTRIBUTION: Everglade Keys of Florida, growing upon decayed wood in hammocks; 
Guadeloupe. 
ILLUSTRATION : Fée, doc. cit. pl. 29, f. 3. 
2. Actinostachys Pennula (Sw.) Hook. Gen. Fil. A/. 217, A. 1842. 
Schizaea Pennula Sw. Syn. Fil. 150, 379, 1806. 
Schizaea trilaterals Schkubr, Krypt. Gew. 1: 137. 1809. 
Acrostichum Pennula Poir. in Lam. Encyc. Suppl. 1: 125. 1810. 
Actinosiachys trilateralis J. Smith, Lond. Jour. Bot. 1: 202. 1842. 
Fronds numerous, closely fasciculate from a small ascending rhizome, stout, 25-50 cm. 
long, strongly triquetrous toward the dark castaneous base and usually buried a distance of 
