30 NORTH AMERICAN FLORA [VoLtumE 17 
sheathing spathe, which splits at maturity. Flowers dioecious, arranged in 2 rows on the 
spadix, each flower covered by a hyaline envelope. Perianth none. Staminate flower of a 
single, sessile, 1-celled anther. Pistillate flower of sessile carpels. Style short, crowned by 
2 capillary stigmas. Mature fruit coriaceous, indehiscent, crowned by the short style at the 
apex and deeply cordate-sagittate at the base. Seeds membranous, globose, not sulcate. 
Type species, Phyllospadix Scouleri Hook. 
Leaves flat, at least 2 mm. wide. 1. P. Scoulert. 
Leaves complicate, 1.5 mm. wide or less. 2. PB. Torreyi. 
1. Phyllospadix Scouleri Hook. Fl. Bor. Am. 2: 171. 1838. 
Phyllospadix serrulatus Rupr.; Aschers. Linnaea 35: 169. 1867. 
Stem winged, mostly simple, 1-4 dm. long ; leaves all submerged, linear, flat 0.5-2 m. 
long, 2-4 mm. wide, entire, rarely serrulate, obtuse at the apex, sheathing at the base; 
primary nerves 3, with usually several intermediary secondary ones ; inflorescence a spadix, 
2-4 cm. long ; spathe prominently winged, half enclosing the flowers ; peduncles basal, sim- 
ple, solitary or in 2’s with sheathing basal bracts, 1-5 cm. long ; staminate flower unknown ; 
pistillate flower of sessile ovaries in 2 rows; style short, crowned with 2 slender stigmas ; 
mature fruit ovate, 2~3 mm. long, 1-seeded, at the apex beaked with the persistent style 
and deeply cordate-sagittate at the base, with often 2 wings projecting downwards from the 
face of the fruit; seed globose, the testa thick. Numerous sterile flowers occur with stami- 
nodia-like appendages. 
TYPE LOCALITY: Dundas Island, Columbia River. 
DISTRIBUTION: Pacific coast from British Columbia to California. 
ILLUSTRATIONS: Hook. loc, cit. pl. 186; Mem. Torrey Club 3?: £1. 73, 74, f. 1-2; E. & P. Nat. 
PA, 21: f. 157, 158. 
2. Phyllospadix Torreyi S. Wats. Proc. Am. Acad. 14: 303. 1879. 
Stems simple or branched, flat, 3 dm. or more long; leaves all submerged, linear, flat 
when very young, in maturity always complicate or truly terete, 0.8-2 m. long, 0.7-1.5 mm. 
wide, the apex obtuse or acute, the base sheathing; primary nerves 1-3, or sometimes the 
leaf-blades nerveless ; inflorescence a curved or straight spadix, the spathe almost enclos- 
ing the flowers and splitting after anthesis; peduncles cauline, mostly branched, in 2’s or 
3’s, mostly 3-14 cm. long, and with numerous fugacious bracts, the bracts scarious, obtuse 
or acute, 1-3.5 cm. long; staminate flowers numerous, in 2 rows, consisting of sessile, 1- 
celled anthers; pistillate flowers of sessile ovaries; mature fruit 1-seeded, flask-shaped, 
2-3 mm. long, beaked by the persistent style at the apex, deeply cordate-sagittate at the 
base, with 2 projecting wings, and sometimes winged on the back ; seeds of a reddish tint. 
Numerous sterile flowers occur. 
TYPE LOCALITY: Santa Barbara, California. 
DISTRIBUTION : Along the coast “of California. 
ILLUSTRATIONS: Mem. Torrey Club 3?: £1. 72, 74, f. 3-5, Engler, PAatwenreicli qu: f. 71. 
