240 NORTH AMERICAN FLORA [VoLUME 22 
Style not articulated to the ovary; hypanthium 
more or less urn-shaped, contracted at the 
mouth. 
Styles nearly basal ; inflorescence cymose ; hy- 
panthium loosely investing the achenes. Tribe 8. ALCHEMILLEAE. 
Styles terminal; inflorescence spicate, race- 
mose, or paniculate ; hypanthium indurate, 
closely investing the achenes in fruit. 
Styles perfectly terminal and straight, or 
slightly bent to one side; herbs with 
spicate or racemose inflorescence, Tribe 9. SANGUISORBEAE. 
Styles inserted at one side of the cushion- 
covered top of the ovary, doubly bent; 
shrubs with paniculate inflorescence. Tribe 10. ADENOSTOMATEAE. 
Disc at the mouth of the hypanthium produced intoa 
cylindric tube, separating the stamens from the 
pistils ; shrubs with opposite leaves and branches. Tribe 11. COLEOGYNEAE. 
g. Seeds inserted at the proximal end of the ovary, 7. é., 
perfectly basal ; radicle inferior. 
Styles wholly deciduous. Tribe 12. COLURIEAE. 
Styles partly or wholly persistent. 
Hypanthium hemispheric, campanulate or turbi- 
nate, persistent. Tribe 13. DRYADEAE. 
Hypanthium salver-shaped, the limb deciduous, 
the tube persistent and closely investing the 
fruit; shrubs. Tribe 14. CERCOCARPEAE. 
8. Fruits of more or less fleshy drupelets; ovules 2, collateral. 
Stamens few or many, inserted on the margin of the flat or 
saucer-shaped hypanthium ; cotyledons flat. 
Pistils at least as many as the ‘sepals, usually numerous ; 
petals present at least in the staminiferous flowers. Tribe 15. RUBEAE. 
Pistils 2-4, fewer than the sepals ; petals in ours none. Tribe 16. KERRIEAE. 
Stamens 15, the 5 antisepalous ones inserted in the tube of 
the campanulate hypanthium ; cotyledons convolute. Tribe 17. OSMARONIEAE. 
II. Carpels enclosed in the hypanthium, which becomes fleshy in 
fruit. Tribe 18. ROSEAE. 
Tribe 1. NEILLIEAE. Shrubs, with alternate, toothed, and usually more 
or less lobed leaves, and deciduous stipules. Inflorescence racemose or corym- 
bose, or rarely paniculate. Hypanthium from hemispheric to deeply campanu- 
late Cin Vez//ia). Disk more or less developed, bearing many stamens; anthers 
didymous. Pistils 1-5; if more than one, more or less united at the base; 
ovules 1-several. Seeds pear-shaped, with a bony shining coat. 
1. OPULASTER Medic. Pfl. Anat. 109. 1799. 
Spiraea § Physocarpos Camb. Ann. Sci. Nat. 1: 239. 1824. 
Icotorus Raf, Bull. Bot. Seringe 216. 1830. 
Epicostorus Raf. Atl, Jour.1: 144. 1832. 
Physocarpa Raf. New F1.3: 73. 1838. Not for ee Necker, 1790. 
Physocarpus Maxim. Acta Hort. Petrop. 6: 219. 187 
Shrubs with exfoliating bark. Flowers in sie corymbs. Leaves alternate, 3-5- 
ribbed, more or less lobed and usually with more or less stellate hairs. Hypanthium hemi- 
spheric or nearly so. Sepals 5, persistent, stellate-tomentulose at least within. Petals 5, 
white or rarely pinkish, spreading. Stamens 20-40, on a disk clothing the mouth of the 
hypanthium ; filaments long, slender, filiform; anthers didymous. Pistils 1-5, more or less 
united at the base; styles filiform, terminal; stigmas capitate; ovules 2-4, the uppermost 
pendulous. Follicles more or less inflated, opening along both sutures; seeds 2-4, most 
commonly 2, obliquely pear-shaped, shining with a bony coat; embryo straight; endo- 
sperm copious. 
Type species, Spiraea opulifolia Ly. 
Carpels 3-5, united only at the base, turgid. 
Mature carpels glabrous, ovate, usually 5. 
Caruncle of the seeds lateral, about one-third as long as the seed ; folli- 
cles usually twice as long ‘as the sepals ; leaves of the sterile shoots 
usually longer than broad, 3-lobed 1. O. opulifolius. 
Caruncle short, almost terminal ; follicles not more than half longer 
rags sepals ; leaves of the sterile shoots scarcely longer than 
road. 
