68 NORTH AMERICAN FLORA [VoLUME 9 
40. GRIFOLA (Micheli) S. F. Gray, Nat. Arr. Brit. Pl. 1: 643. 1821. 
Polypilus Karst. Rev. Myc. 39: 17. 1881. 
Meripilus Karst. Bidr. Finl. Nat. Folk 37: 33. 1882. 
Cladomeris Quél. Ench. Fung. 167. 1886. 
Hymenophore large, annual, stipitate, compound, intricately branched or lobed, 
humus-loving or epixylous, rarely terrestrial, usually found at the base of a tree-trunk ; 
surface smooth, pallid to gray or brown: context white, fleshy or fleshy-tough, rigid and 
fragile when dry; tubes large, irregular, thin-walled, becoming friable or laciniate with 
age: spores hyaline, smooth, rarely verrucose. 
Type species, Boletus frondosus Dicks. 
Hymenium ochraceous, becoming dirty-yellow with age; plants terrestrial, 
irregularly confluent, olivaceous to greenish-yellow. 
Hymenium at first fuliginous, becoming paler. 
Hymenium white or pallid from the first. a 
Surface of pileus gray or grayish-brown to coffee-colored ; stipe intricately 
branched ; pileoli very numerous and small. 
Pileoli lateral, spatulate or dimidiate. 3. G. frondosa. 
Pileoli centrally attached, circular and umbilicate. 4. G, ramosissima. 
Surface of pileus pallid or alutaceous; stipe not intricately branched, lobes 
usually few in number and comparatively large. , 
Sporophore of immense size, 20-60 cm. in diameter; spores echinulate, 
1. G. poripes. | 
2. G. Sumstinet. 
5. G. Berkeleyt. 
M, 
Sporophore small for the genus, only 8 cm. or less in diameter; spores . 
smooth, ovoid, much smaller. 6. G. fractipes. 
1. Grifola poripes (Fries) Murrill, Bull. Torrey 
Club 31: 335. 1904. 
Polyporus poripes Fries, Nov. Symb. 48. 1851. 
Polyporus flavovirens Berk. & Rav. Ann. Mag. Nat. Hist. II. 12: 431. 1853.—Grevillea 1: 38. 
1872. (Type from South Carolina.) 
Pileus at first simple and centrally stipitate, becoming imbricate-multiplex when fully 
developed, 8-20 cm. in diameter; pileoli soft, fleshy, fragile when dry, circular to flabelli- 
form, pulvinate or depressed to applanate, 5-10 cm. broad, 5-8 mm. thick ; surface sordid- 
yellow, with yellowish-green zones, becoming dull yellowish-green, finely tomentose to 
subglabrous; margin irregular, undulate to lobed, concolorous: context fleshy, very 
fragile when dry, 24 mm. thick, white to yellowish; tubes very decurrent, yellow to 
yellowish-green, 3-5 mm. long, mouths irregular, circular to sinuous, 1-2 to a mm., at first 
milk-white, becoming dirty-yellow, edges thin, fragile, lacerate with age: spores sub- 
globose, smooth, hyaline, 34.5%: stipe central or excentric, pallid, 3-6 cm. long, 1-1.5 cm. 
thick, becoming tubercular and connate-ramose at maturity. 
TYPE LOCALITY : North Carolina. 
HABITAT: On the ground in woods. 
DISTRIBUTION: Eastern United States, west to Missouri. 
ExsiccaTI: Rav. Fungi Car.4: 4; Ellis & Ev. N. Am. Fungi 1689. 
2. Grifola Sumstinei Murrill, Bull. Torrey Club 31: 335. 1904. 
A very large plant resembling G. frondosa in habit and general appearance, but with 
fewer and broader pileoli, darker surface and darker hymeninm. Pileus imbricate-multi- 
plex, 20 X 30 cm.; pileoli flabelliform to spatulate, 6-8 X 6-8 < 0.3-0.5 cm.; surface radiate- 
rugose, finely tomentose, light- to dark-brown ; margin very thin, fissured and strongly in- 
flexed when dry: context white, fibrous, fleshy-tough to almost leathery, 0.3 cm. thick ; 
tubes 0.2 cm. long, 7 to a mm., at first fuliginous, becoming pallid at maturity, polygonal, 
irregular, edges very thin and fragile, becoming lacerate: spores globose, smooth, hyaline, 
thin-walled, copious, 5 #: stipe tubercular, woody, blackish below, connate-ramose, lighter- 
colored, passing insensibly into the pileoli above. 
TYPE LOCALITY: Pennsylvania. 
HapitaT: About old stumps and trunks of deciduous trees. 
DISTRIBUTION : New York, Pennsylvania, Virginia, Ohio, and Louisiana, 
