216 NORTH AMERICAN FLORA [VorumE 21 
long, 1.5-3 em. wide, cordate at the base and slightly unequal, acute or attenuate at the apex, 
minutely puberulent when young but soon gabrate; flowers in 3-6-flowered umbels on long 
slender peduncles, the pedicels 1-2 cm. long, short-villous or rarely glabrate; bracts linear, 
2-3 mm. long, green, short-villous, caducous; perianth 7-10 mm. long, 10-15 mm. broad, 
white, short-villous outside, at least below; stamens 2, exserted; fruit narrowly clavate-oblong, 
5-10 mm. long, 2 mm. thick, dark-green, glabrous, bearing numerous glands arranged in 
3 or 4 transverse bands. 
TYPE LOCALITY: San Pablo, Lower California. 
DistRIBUTION: Southern Lower California. 
19. ANULOCAULIS Standley, Contr. U. S. Nat. Herb. 12: 374. 
1909. 
Tall erect perennial herbs, more or less pubescent, branched, the internodes usually 
each with a viscid ring. Leaves opposite, petiolate, few, borne near the base of the stem, the 
blades broad, coriaceous, glandular-dentate or denticulate. Flowers perfect, bracteate, um- 
bellate or in axillary glornerules or racemose, the inflorescence ample, much branched, the 
bracts small, scarious or coriaceous; perianth funnelform, the tube elongate, constricted above 
the ovary, the limb campanulate or subrotate, 5-lobed, induplicate-plicate. Stamens 3 or 5, 
exserted; filaments unequal, filiform, connate at the base; anthers didymous. Style filiform, 
the stigma peltate. Amnthocarp coriaceous, biturbinate, 10-striate, glabrous, in one species 
(probably in all) developing at maturity a broad median horizontal wing. Seed with its 
thin testa adherent to the pericarp; embryo uncinate, the broad cotyledons enclosing the 
copious endosperm; radicle elongate, descending. 
Type species, Boerhaavia eriosolena A. Gray. 
Perianth glabrous, 2.5-3 cm. long; leaf-blades glabrous or nearly so. 1. A. leiosolenus. 
Perianth villous, 8-10 mm. long; leaf-blades long-villous beneath, at least when 
young. 
Flowers mostly in sessile axillary clusters; perianth abruptly expanded into a 
subrotate limb. 2. A. eriosolenus. 
Flowers in pedunculate terminal umbels; perianth gradually dilated into a 
campanulate limb. 3. A. annulatus. 
1. Anulocaulis leiosolenus (Torr.) Standley, Contr. U. S. Nat. Herb. 
12: 375. 1909. 
Boerhaavia leiosolena Torr. Bot. Mex. Bound. Surv. 172. 1859. 
Acleisanthes nummularia M. E. Jones, Contr. West. Bot. 10: 43. 1902. 
Boerhaavia nummularia M. E. Jones; Prain, Ind. Kew. Suppl. 4: 27. 1913. 
Plants erect, 6-10 dm. high, sparsely branched below, the brancies stout, glaucous, 
glabrous; leaves 1 or 2 pairs at the base of the stem, the petioles stout, 2-11 cm. long, the 
blades reniform to broadly oval, rhombic-orbicular, or ovate-oval, 3.5-15 cm. long, 3.5-12.5 
cm. wide, deeply cordate to truncate at the base, broadly rounded at the apex, rather 
coarsely sinuate, thick-coriaceous, yellowish-green on the upper surface and when young 
tomentulose, beneath glaucescent and tomentulose or villous when young but early glabrate 
and gland-dotted; inflorescence much branched, the branches stout, naked, the flowers fascicu- 
late or in dense spikes, short-pedicellate; bracts minute, ovate, coriaceous, glabrous; perianth 
yellowish-green, 2.5—-3 cm. long, glabrous, gradually dilated above into a narrow limb; stamens 
5, short-exserted, fruit biturbinate, 5-6 mm. long, glaucous, in age developing at the middle a 
rigid horizontal wing 6-7 mm. in diameter; seed biturbinate, 3 mm. long, pale-brown. 
Type LocaLity: In gypseous soil, Great Canyon of the Rio Grande, 70 miles below El Paso, 
Texas. 
DISTRIBUTION: In strongly alkaline soil, western Texas to Arizona and Nevada. 
2. Anulocaulis eriosolenus (A. Gray) Standley, Contr. U. S. Nat. 
Herb. 12: 375. 1909. 
Boerheavia eriosolena A. Gray, Am. Jour. Sci. If, 15: 322. 1853. 
Plants erect, 2-10 dm. high, sparsely branched below, the branches very stout, glabrous; 
petioles stout, 1-4 cm. long; leaf-blades orbicular-oval to ovate-oval, 2.2-10 em. long, 2-7 cm. 
