Manual de Plantas de Costa Rica | The physical environment | 89 
families pollinated by dynastine scarab beetles at the La Selva Biological Station often 
flower during the rainier months. At the same site, shrubs and small trees specialized 
in colonizing new light gaps appear to begin flowering whenever they reach a certain 
critical size, regardless of season, and to continue flowering until they are shaded out. 
Flowering of many other species is apparently linked to day-length, rather than rainfall. 
For additional information on phenological patterns in Costa Rican plants, see espe- 
cially Fournier O. & Salas (1966), Janzen (1967), Daubenmire (1972), Frankie et al. 
(1974), Opler (1980), Bullock (1981), Koptur et al. (1988), and Bawa et al. (2003). 
See Holdridge et al. (1971), Coen (1983), and, especially, G6mez P. (1986) and 
Herrera (1985) for further information on the global factors responsible for Costa Rican 
climate and weather, classification and formal descriptions of Costa Rican climatic 
groups, local weather phenomena, and copious weather data for individual sites, in- 
cluding tables, graphs, and maps. 
