Manual de Plantas de Costa Rica 229 
area worthy of, and in need of, much more exploration. These trips have resulted in a 
great number of additions to the flora, including Agouticarpa williamsii, Anthopterus 
costaricensis*, A. revolutus, Anthurium al- 
ticola, A. antonioanum, A. bittneri*, A. cu- 
cullispathum, A. hornitense, A. longistipi- 
tatum, Byrsonima herrerae, Calyptrogyne 
herrerae*, Campyloneurum magnificum, 
Cavendishia chiriquensis, C. limonensis*, 
Chamaedorea anemophila, C. rosibeliae*, 
Coussarea duplex*, Dacryodes sp., Daph- 
nopsis mori, Disterigma trimerum, Elaph- 
oglossum bittneri*, E. nigrosquama*, Geo- 
noma talamancana*, Licania  jefensis, 
Macoubea mesoamericana, Marila jefensis, 
Metteniusa tessmanniana, Micropholis guy- 
anensis, Monolena primuliflora, Neurolepis 
pittieri, Nyssa talamancana, Ocotea rufes- 
cens*, Peperomia ocumarana, P. ursina*, 
P. villarrealii, Perebea guianensis, Pouteria 
ramiflora, Pterozonium brevifrons, Rhodos- 
patha moritziana, Rhynchospora cabecar- 
ae*, Senecio angustiradiatus, Stenosperma- 
tion pteropus*, Tetranema gamboanum*, Themistoclesia horquetensis, Triolena pumila, 
and 7: spicata. The majority of these new records are from Panama or South America. 
Neurolepis pittieri 
Lowlands and minor mountain ranges 
The Llanura de Los Guatusos, where few protected areas exist, has been heavily de- 
graded by agricultural and lumbering activities, but even so, it accounts for a number 
of new records of great interest for the country, such as Acoelorraphe wrightii, Chris- 
tiana africana, Hirtella guatemalensis, Inga capitata, Miconia chamissois, M. tomen- 
tosa, Solanum ipomoea, Tetrapterys mucronata, and Zygia conzattii. A few of these 
species give a hint of the relationship between this area and the Caribbean slope of 
northern Central America. Among the new species that have been found here are 
Gamanthera herrerae*, Inga canonegrensis*, and Sclerolobium costaricense. 
The Llanuras de San Carlos and Sarapiqui have also been modified by human ac- 
tivity, but their combination of flatlands and hills harbors a diverse flora that has con- 
tributed considerably to our list of new discoveries. A few of these are Caryodendron 
angustifolium, Cayaponia sessiliflora, Chorigyne cylindrica, Conchocarpus nicaragu- 
ensis, Doliocarpus dasyanthus, D. foreroi, Elvasia elvasioides, Henriettella odorata, 
Ipomoea eremnobrocha, Licania affinis, Malanea erecta, Miconia sparrei, Pouteria 
