41 
worked over the two areas, succinctly summarizes the whole relation of 
both floras. 
This Malay-Papuan influence is the determining factor extending to the 
South Sea Islands, while the Moluccas show Malay with Philippine ground- 
types and a strong Papuan influence (28, i. 14, xx), and Schlechter describes 
New Guinea as the centre of distribution of an endemic Orchid flora ( Aus- 
gangscentrum einer eigener Orchideenflora ”) (23, i. 14, xx) as evidenced 
by Corysanthes, supposed to be Australian, but now truly Papuan, 13 species 
being known from N.E. New Guinea alone. The presence of this genus in 
Australia, New Caledonia, Samoa, Java, Philippines, and the Himalayas 
suggests radiation from a Papuan centre of development (23, 14, xxiii). 
This overwhelming Papuan influence is amply demonstrated in many 
other families, which not only show an actual numerical predominance in 
species, but also a greater range of specific differentiation than is known 
from elsewhere : for example, Libocedrus (4), Drimys (19), Myrtus (6) ; of 
Pandanus and Freyeinetia, 1 have never seen such a wide range in form ; 
while in Rhododendron, Vaccinium, Styphelia, the numerical predominance 
along with the great morphological range of form is unsurpassed in any 
other region, except perhaps, in the case of the first, Central China. 
Further interesting proof of the soundness of this point of view is afforded 
by comparison of the Australian and New Guinea representatives of the 
same genera, the Papuan forms showing decidedly the oldest types. 
In the present collection perhaps Trimenia is the most striking case in 
point. 7. weinmanniefolia Seem., described in 1852 from Fiji, a dicecious 
plant, remained the type of a supposed monotypic Polynesian genus, closely 
allied to Piptocalye Moorei, also dioecious, ranking as a monotypic Australian, 
one, till Ridley described T. papuana from Mt. Carstensz in 1916. 7. arfak- 
ensis is included in the present paper. 
Two species of Zrimenia are now known from New Guinea, both 
hermaphrodite, while two new species in a closely allied new genus, Jden- 
burgia, show a syncarpous bilocular ovary, proving not only that the dicecious 
habit of the two isolated outliers of this order is probably derived, but also 
that the systematic position of Zrimenia and Piptocalya in the apocarpous 
Monimiacez is untenable, necessitating the new order Trimeniacez. 
An equally convincing example is that of Pullea, a genus established by 
Schlechter in Saxifragacee to include two plants with inferior ovary from 
N.E. and N.W. New Guinea respectively, to which P. papuana is now 
added, with a N. Queensland species, P. Stutzeri = Callicoma Stutzeri 
F. Muell., first distributed as Stutzeria by him, but afterwards included in 
Callicoma. Pullea, therefore, now includes three distinct Papuan species 
and one N, Australian, but, had the latter plant first been accorded proper 
generic position, the former would have been cited as evidence of a wave of 
Australian immigration into New Guinea, 
