130 STRUCTURE OF THE OVARY OF MARLEA, 
Alungiee ; and among hermaphrodite epigynous Orders that have been 
compared with this Order, this bifid stigma exists only in a part of the 
Order Cornacee, among which the stigma of Cornus alba may be con- 
sidered analogous to those of Marlea. As in Marlea, there are four 
lobes, the lobes being more united; an open canal extends through 
the style down into the disc, though it does not enter the ovary ; and 
the position of the cells of the ovary being compared with that of the 
lobes of the stigma, they are found to correspond with the fissures of 
the stigma, nearly, though not so precisely, as in Marlea. The cells 
are, however, constantly opposite the two angles in the canal at the 
base of the style, which are continued from two opposite fissures in the 
stigma, the other two having become obsolete as in Marlea (Vide Pl. 
V. B. f. 1, 2, 8, 4 and 5). : 
But in Cornus sanguinea the cells of the ovary are opposite two 
lobes of the stigma, which are, moreover, unequal (Vide Pl. V. C. f. 1); 
yet the similarity between this species and C. alba is such, that the 
structure can hardly be thought to differ; and that it is the same as 
C. alba, is vendered probable by the internal processes which descend 
from the lesser lobes of the stigma, not disappearing in the style below 
(as when stigmas represent abortive carpels), but increasing so as to 
overlap each other (Vide Pl. V. C. f. 1, 2 and 3), This imbrieation 
of opposite lobes does not take place in Marlea and in C. alba, they 
only lie in contact (Vide Pl. V. B. f. 2), which may be sufficient to 
account for the cells of the ovary of C. sanguinea not having the same 
relation to the lobes of the stigma as in C. alba. The number also of 
the lobes of the stigma of C. sanguinea is generally double the number 
of the carpels, or nearly so, when there is an increase of carpels in the 
ovary ; thus, an ovary having four cells had a stigma with eight or 
nine lobes, and another with six cells had a stigma with eleven lobes 
(Vide Pl. V. C. f. 4 and 5). À 
Another indication of the affinity existing between Marlea and 
Cornus, and also between Alangium and Cornus, is furnished by the 
changes which occur in the structure of Cornus sanguinea. 1 have 
found, when cultivated in a nursery, the number of petals and stamens 
_ frequently increased to 5, 6, 7, or more, the greatest increase observed 
being 8 petals and 10 stamens, the ovary of the flower being four- 
celled ; the carpels; however, sometimes inerease to 7, without any fur- 
ther increase of the petals and stamens. Thus the numbers of the parts 
