LUNDELL: THE GENUS PARATHESIS 177 
69. PARATHESIS ADENANTHERA (Miq.) Hook.f., ex Mez, Pflanzenreich IV. 
236: 179. 1902. Fig. 66 and Plate 32. 
Ardisia adenanthera Migq., Fl. Brasil. 10: 285. 1856. 
Tinus adenanthera (Miq.) O. Ktze, Rev. Gen. 2: 974. 1891. 
Ardisia ferruginea H.B.K. var.? macrophylla Benth., Pl. Hartweg. 217. 1846 ; Benth. & Hook. 
f., Gen. Pl. 2: 645. 1876. 
Parathesis macrophylla (Benth.) Britton, ex Rusby, Mem. Torrey Club 4: 217. 1895, not 
P. macrophylla Rusby, ex Mez, Pflanzenreich IV. 236: 180. 1902. 
Parathesis rubella Moldenke, Phytologia 1: 11. 1933. 
Shrub or tree, up to 6 m. high, branchlets rather stout, at first ferruginous- 
tomentose with appressed stellate hairs; leaves with petioles 1-2 em. long; leaf 
blades elliptic, oblong-elliptic or obovate-elliptic, 10.5-28 em. long, 5-10 em. 
wide, apex subabruptly acuminate, base acute to acuminate, membranaceous, 
irregularly undulate and crenulate, sometimes appearing subentire, at first 
pubescent on lower surface with small ferruginous appressed stellate hairs, 
glabrescent, costa slightly impressed above, elevated and conspicuous on under 
surface, the primary lateral veins slender; inflorescences axillary, usually very 
large, open, lax, tripinnately paniculate, 15-33 cm. long, the peduncle and 
primary and secondary branches long, minutely ferruginous-puberulent, often 
densely so; pedicels slender, 4-7 mm. long, in fruit up to 13 mm. long; flowers 
subcorymbose, densely and minutely ferruginous-puberulent, punctate with 
reddish glands, 5-7 mm. long at anthesis; sepals narrowly triangular, 1-1.5 mm. 
long, acuminate, minutely puberulent, densely reddish-punctate; petals narrowly 
triangular-lanceolate, 5-7 mm. long, papillose-tomentulose on inner surface, 
densely reddish-punctate in lines; stamens 2.5-4 mm. long; filaments slender, 
2-3 mm. long, pubescent with short gland-tipped hairs; anthers erect, small, 
lanceolate, 0.8-1.5 mm. long, apiculate, with several small reddish glands dor- 
sally, dorsifixed about two-fifths above base; ovary ovoid, ferruginous-tomentu- 
lose apically and at base of style with short shaggy hairs; style 3-4.5 mm. long; 
placenta depressed-globose, apiculate; ovules 5-8, uniseriate, enclosed; fruits 
subglobose, up to 16 mm. diam. when ripe, ferruginous-puberulent at first, 
glabrescent, costate. 
COSTA RICA.—Yerba Buena, northeast of San Isidro, Provincia de Heredia, wet forest, 
alt. about 2000 m., Feb. 22, 28, 1926, Paul C. Standley & Juvenal Valerio 49856 (US), shrub 
10 ft., fruit pale red; San Jose, Cerro Gallito, hasta 2000 m., Feb. 3, 1935, Manuel Valerio 
1190 (F). 
COLOMBIA.—Prov. Bogota, Hacienda de Palmar, village of Guadua, Karl Theodor 
Hartweg s.n. (K, type of Ardisia ferruginia H.B.K. var.? macrophylla Benth.), a shrub 6 ft. 
high; Valparaiso, vicinity of Medellin, July 22, 1928, Rafael A. Toro 1363 (NY); Dept. Boyaca, 
region of Mt. Chapon, northwest of Bogota, stream-side in high thick forest, alt. 6000 ft., 
July 31, 1932, A. E. Lawrence 372 (NY, type of P. rubella; A, BM, F, G, GH, MO, §, isotypes), 
tree 15-20 ft., 8-12 in., fls. pink; Dept. del Valle, Cordillera Occidental, vertiente occidental, 
Hoya del Rio Sanquinini, lado izquierdo, La Laguna, bosques, alt. 1250-1400 m., Dec. 10-20, 
1943, Jose Cuatrecasas 15426 (F), arbolito; Hoya del Rio Cali, Pichinde, entre La Marina y 
La Margarita, potreros y residuos de Monte, alt. 2120-2260 m., Nov. 4, 1944, Jose Cuatrecasas 
18600 (F, G), arbolito, 6 m., drupas rojas, brillantes, esferico aplastadas 16 mm. diam.; Valle, 
La Cumbre, quebrada La Ventura, alt. 1650 m., Apr. 9, 1947, Lorenzo Uribe Uribe 1552 (F); 
without locality, 1760-1808, Jose Celestino Mutis 474 (US); Justin Goudot s.n. (G, K, P). 
PERU.—Cuchero, E. F. Poeppig 1667 (F, GH, P, isotypes); La Merced, alt. ca. 2000 ft., 
