OF MUTANT CHARACTERS. 



211 



ANTLERED fyf). 



(Plate 9.) 



ORIGIN OF ANTLERED. 



The character "antlered" was found by Morgan about September 

 1912, and a pure stock was secured. It seems to have originated in 

 an experiment involving vestigial. 



DESCRIPTION OF ANTLERED. 



The wings of antlered flies are on the average longer than those of 

 strap, often, indeed, being full length. The wing is also broader in 

 the distal portion, so that sometimes it can scarcely be distinguished 

 in form from a rather extreme beaded. Like strap, too, the wings are 

 held out at rather wide angles (about 30 from the axis of the body). 

 A unique feature of antlered is that the long type of wing is quite often 

 folded at the tip (see fig. 2, plate 9). 



INHERITANCE OF ANTLERED. 



Rather extensive breeding and selection experiments were carried 

 out on this wing, the records of which have been lost. The results, 

 however, were in agreement with some later data, which may be 

 given. Antlered males out-crossed to wild females gave wild-type 

 FI offspring. Seventeen mass-cultures of the FI flies were bred, giving 

 in F 2 a total of 5,234 flies, of which 1,036 or 19.8 per cent were antlered 

 (August 1915; Morgan, 1915, p. 10). That is, antlered is a simple 

 recessive to wild, and the F 2 ratio was aberrant because of the crowding 



TABLE 57. PI, antlered cf cf X vestigial 9 9 . 



vestigial ? cf cT . 



vestigial 9 9 + 



that always takes place in mass-cultures. The antlered did not split 

 up in F 2 , which shows that antlered is not vestigial plus a modifier 

 unless that modifier is in the second chromosome linked so closely to 

 vestigial that no appreciable crossing-over occurred. 



When antlered males were crossed to vestigial females the FI flies 

 were not wild- type. They are recorded as being all vestigial (Feb- 



