ADDENDUM TO SPONGES 



167 



about 10 mm. in height by 5 mm. in breadth. The openings of the 

 canal system are confined to the upper surface (Fig. 97, A; cf. Tentwium, 

 Fig. 31). 



The skeleton of Astrosclera is composed, not of spicules, which are 

 entirely wanting, but of calcareous spherules, which arise in cells of the 

 dermal layer near the upper surface. Each spherule is deposited within 

 a single cell, and is from the first composed of radially arranged crystalline 

 fibres. Its form is at first spherical, but by further increase in size 

 adjacent spherules come into contact, and the interspaces between them 

 become completely filled in by continued deposition of the calcareous 



FIG. 97. 



Astrosclera Willeyana, Lister. A, the sponge magnified about three diameters ; p.s., ripper 

 surface carrying the openings of the canal system ; 6, base of attachment. B, section of the 

 skeleton ; sph., spherules ; c, canals. (Drawings by Mr. J. J. Lister.) 



/ 



substance, to the exclusion finally of the soft parts. The spherules thus 

 acquire a polyhedral form (Fig. 97, B, sph.), and by their union build up 

 a solid calcareous skeleton without any admixture of soft parts, but 

 traversed by canals in which are lodged the soft tissues and the canal 

 system of the sponge (Fig. 97, B, c). In the basal (older) part of the 

 sponge the canals became obliterated, apparently by extension inwards of 

 the spherules forming their wall ; just as in pedunculate sponges the 

 canal system is wanting in the stalk. 



The spherules are composed of aragonite, and contain an organic 

 basis which has the same staining reactions as that of the spicules of 

 Calcarea. 



The canal system is of a leuconoid type with small spherical ciliated 

 chambers opening into branched canals. There is no large central 

 gastral cavity, but a number of excurrent canals [which perhaps represent 

 gastral cavities reduced in size] run vertically upwards to open on the 

 upper surface, alternating with in current canals similar in appearance and 

 arrangement. Some ova and larvae were observed, the latter apparently 

 of a parenchymula type. 



The affinities of this curious organism are very doubtful. It is un- 

 questionably a sponge, and the material of its skeleton is carbonate of 



