154 Spurious A llelonwrphism [CH. 



The hooded types may have a great diversity of colours, 

 and fixed hooded varieties now exist in the purple, blue, 

 red, pink, cream and other classes. It is nevertheless a 

 remarkable fact that, so far as I am aware, none of the 

 regular bicolour varieties ever have a really hooded standard. 

 There is for instance no hooded type having the colour of 

 the original purple, with its chocolate-purple standard and 

 blue wings, nor can Painted Lady with standard red and 

 wings nearly white be produced in a hooded shape. On 

 the contrary the hooded types always have the standard and 

 wings more nearly alike in colour, and there is the clearest 

 evidence that in families (F 2 and later generations) which 

 contain original bicolour purples as well as hooded types, 

 the hooded types corresponding to them are of the uni- 

 colorous kind known as " Duke of Westminster*." 



From these facts it is evident that there is here some 

 interdependence between the colour of the flower and its 

 form. This interdependence is of course somatic, but as 

 will be seen there is also a gametic connection between the 

 phenomena of shape and colour. 



The experiments bearing on these questions originated 

 in a cross between the white, round-pollened Emily Hen- 

 derson and a white, long-pollened hooded type known as 

 Blanche Burpee. The Emily Henderson has an ordinary 

 erect standard with the central notch. 



F l produced from these two is a bicolour purple, with 

 erect standard and long pollen, indistinguishable from the 

 reversionary /^ previously described as the offspring of the 

 long and round whites. F^ from such plants consists of 

 the following types : 



Purple Red White 



bicolour unicolorous bicolour erect hooded 



erect hooded erect 



81 27 36 84 28 



108 112 



27 9 



36 28 



9 7 



* Similarly if the bicolour purples with dark wings are present in the 

 class with an erect standard, they are represented by "Duke of Sutherland" 

 in the hooded class, viz. a deep unicolorous purple. 



