STUDIES ON THE SECONDARY SEXUAL 



CHARACTERS OF CRAYFISHES. V. 



MALES WITH FEMALE 



CHARACTERS. 



C. L. TURNER, 

 ZOOLOGICAL LABORATORY, BELOIT COLLEGE. 



A number of crayfishes belonging to the genus Cambarus in 

 which secondary sexual characters of both sexes were found upon 

 the same individual have been described by Faxon, Hay, Ortman 

 and the writer. All of these specimens with the exception of 

 one described by Hay were females. In some cases there was a 

 full complement of female secondary sexual characters with the 

 addition of one or more male characters and in other cases a 

 female character was lacking while one or more male characters 

 were present. The one exceptional specimen described by Hay 

 was hermaphroditic and possessed both male and female primary 

 and secondary sexual structures. 



Male crayfishes in the genus Cambarus bearing sexual char- 

 acters of females have not been noted before and it is the purpose 

 of this paper to describe two specimens which appeared in recent 

 collections made in southeastern Wisconsin. Both specimens 

 were collected from tributaries of the Rock River near Beloit, 

 Wisconsin, the first in July, 1923, and the second in July, 1924. 



One is a specimen of Cambarus virilis, 65 mm. long and is a 

 second form male normal in every respect except that an oviducal 

 opening exists upon the left third walking leg. An examination 

 of the internal organs shows a normal spermary and normal 

 spermatic ducts. There is no tube of any kind joined to the 

 oviducal opening at the base of the third walking leg. The 

 second specimen is one of C. propinquus, 46 mm. long, and is 

 identical in its peculiarities with the first specimen described. 

 It is a normal second form male with all the normal internal and 

 external structures of the male but bearing in addition an ovi- 

 ducal opening at the base of the third walking leg. As in the 



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