CLOACA AND SPERMATHECA OF HEMIDACTYLIUM. 283 



The glands described here are all confined to the region of (i.e., 

 located in the substance of) the dorsal elevation, but there are in 

 addition two other groups. One group, consisting of four or 

 five tubules on each side, lying dorsal to and opening into the 

 roof of the cloaca, is located entirely caudal to the elevation. This 

 group is separated by a short but definite interval from those 

 intermingled with the spermathecal tubules. The other group, 

 also of four or five pairs, is located in the region of the dorsal 

 expansion, therefore cephalic to the spermatheca. 2 



FIG. 2. Cloaca of Hemidactylium (adult, 7.1 cm.) at a level through the openings 

 of the spermathecal tubules into the lateral walls of the mid-dorsal slit. Com- 

 parison with Fig. i shows that, as the tubules pass cephalicly toward their openings, 

 they also slope ventrally, and now lie within the substance of the dorsal elevation. 

 The elevation is composed of two folds or limbs with the mid-dorsal slit between 

 them. The elevation ends abruptly a few sections cephalic to this level. Above, 

 the mid-dorsal slit is already merging into the dorsal expansion ahead of the 

 elevation. The tubules on the right side end one section cephalic to this level; 

 on the left side, two of the three tubules are seen bending sharply toward the slit. 

 (The difference between the sides is due to slight obliquity of sectioning.) Pigment 

 appears above the cloaca. ( X 30.) 



The epithelium of these glands is emphatically not that of func- 

 tional gland tubules, as seen in the ventral group in adult Gyrino- 

 pJiilus; for the tubules are minute, the lumen barely discernible in 

 most of the tubules, and the cytoplasm in each cell scanty. They 

 closely resemble early stages in the development of glands in 

 young Gyrinophilus females, and are apparently rudimentary or 

 only slightly developed structures. Rudimentary dorsal glands 

 were found in Eurycca (Spelcrpcs) in two groups (Kingsbury 

 '95), one cephalic to and one caudal to the opening of the spenna- 



2 Another adult measuring 5.5 cm. (with ova i.o mm. in August), which has 

 since been examined, shows only the most cephalic of these three groups of glands. 

 They are probably quite variable, as are the rudimentary glands of Gyrinophilus. 



