DIGESTION IN THE STOMACH 



777 



account for the whole of the gastric juice obtained as the result of 

 a meal. If an animal provided with two gastric fistulsB, one into 

 a diverticulum and the other into the main stomach, has both its 

 vagi divided, it is found that the introduction of meat into the large 

 stomach is followed, after a period of twenty to forty-five minutes, 

 by the appearance of a secretion of gastric juice from the small 

 stomach. Moreover, when an animal is given a normal meal and 

 is allowed to swallow the food after mastication, the total amount of 

 gastric juice obtained is greater than that produced by the sham 

 feeding alone, and the flow is of longer duration. In fact, we may 

 say that the gastric juice secreted in response to a normal meal 

 consists of two parts, viz. : (1) a large amount, the secretion of which 

 begins within five minutes of the taking of the food and is determined 

 by the reflex nervous mechanism described above ; and (2) a smaller 

 portion, the secretion of which is excited by the presence of the 

 food in the stomach. This combined character of the gastric juice 

 produced by a normal meal is shown in the following Table (Pawlow) : 



SECRETION OF GASTRIC JUICED 



In the first column is given the result of a normal meal on the 

 secretion from the gastric diverticulum. In the second column 

 are given the amount and digestive power of the juice which is excited 

 by the direct introduction of 150 grm. of meat into the large stomach 

 of the animal, care being taken not to excite in any way the nervous 

 reflex mechanism. In the third column are given the amount and 

 digestive power of the juice which is evoked by a sham meal of 

 200 grm. of meat. In the fourth column is given the sum of the last 

 two experiments. It will be seen that the total effect of the sham 

 meal plus the direct introduction of meat into the stomach is almost 

 identical with the secretion obtained when the food is taken in a 

 normal way and allowed to pass through the oesophagus into the 

 stomach. 



The second phase of the gastric secretion cannot be ascribed to 



