HARPALIN^E 47 



of Dr. Horn, by whom it appears to have been first brought to 

 notice and was employed by that author merely in the separation 

 of genera which seemed to be otherwise closely allied. 



In the Harpalinse there are numerous confusing parallelisms of 

 structure, appearing in genera evidently widely separated in the 

 present stage of evolution and probably reversional in nature, or 

 indicative of like conditions of environment during the progress 

 of evolutionary changes, such for instance as the occurrence of 

 series of setigerous substrial punctures in such widely separated 

 types as Stenomorphus, Selenophorus and Philodes, or the occurrence 

 of the trifid anterior tibial spur in various not closely related genera 

 within the tribe Anisodactylini, also occurring in the Amarid genus 

 TricEna and so of no special significance in the estimation of generic 

 relationship. Again, the presence of the enlarged basal joint of 

 the anterior tarsi, especially of the female, characterizing the highly 

 specialized genera Stenomorphus and Gynandropus, as well as 

 Gynandrotarsus harpaloides of the Anisodactylini, and, finally, the 

 occurrence of the rare seta at the hind angles of the prothorax in 

 the indubitable Anisodactylid Diachromus and in Trichocellus of 

 the Acupalpini. Dicheirotrichus is somewhat intermediate between 

 these two genera, tending to unite the two divisions based upon the 

 labial palpi and, as significant in this respect, there seem to be only 

 three long anterior setae on the second labio-palpal joint; the male 

 tarsi, also, are not so purely Anisodactylid as in Diachromus, 

 although the rather even distribution of the long hairs of the soles 

 would seem to betoken somewhat more of an Anisodactylid than 

 Acupalpid affinity, the general habitus of the body, however, being 

 evidently more nearly that of the Acupalpini, to which tribe it is 

 here attached. 



The various tribes of the Harpalinse, so far as represented by 

 material accessible to me, may be defined as follows. The exotic 

 groups, whether tribes or genera, in this and all subsequent tabular 

 statements, are indicated by a prefixed asterisk as usual : 



Second joint of the labial palpi plurisetose in front 2 



Second joint bisetose in front 6 



2 Middle tarsi (cf ) not modified in vestiture beneath and undilated, 

 the anterior sometimes so modified, however, as in Geopinus; ab- 

 domen generally with accessory setae as in a large division of Har- 

 palus; basal angles of the prothorax generally acutely rectangular 

 or acutely subeverted, the prosternum short DAPTINI 



