120 



Bottcher ( '69. 72) disagreed with Hensen, though he has con- 

 firmed, as did Middendorp ('67), the striking inward spreading 

 of the base of the inner pillar cell. 



Gottstein] ( 72) held that the inner pillar cell does not move 

 inward, but that the increase in the length of the labium tym- 

 panicum may explain the peculiar approach of the habenula 

 perforata to the arch of Corti. 



Retzius ('84) agreed in general with Hensen 's assertion that 

 in the course of development the surface of the sense organ 

 comes to lie under the basal surface of the membrana tectoria. 

 He thought that this change of position is brought about "weni- 

 ger in dem Verhalten der Pfeilerzellen, sondern vor allem in 

 dem starken Wachstum der Deitersschen Zellen und der von 

 aussen andriickenden Hensenschen Stiitzzellen, ' and that, 

 further, "vielleicht die Membrana tectoria selbst durch eigenes 

 Wachstum und durch Vergrosserung des Limbus mit seinem 

 Vorspriingen" contributes to this. 



Held ('09) agrees with Hensen on the whole. 



Prentiss ('13, p. 450) denies the wandering of the spiral organ 

 as follows: ''There is no necessity for, and my preparations 

 afford no proof of, an inward shifting of the spiral organ and 

 a consequent displacement of the membrana tectoria " 



Hardesty ('15, pp. 60 and 61) discussed the relative position 

 of the spiral organ with reference to the basal surface of the 

 tectorial membrane and says " the developed spiral organ acquires 

 its position well under the basal surface of the tectorial membrane 

 almost entirely by being carried axisward during the completion 

 of the membrane." "In the apical turn, where these changes 

 are greatest, the hair cells of the organ may be carried axisward 

 a distance nearly half the width of the membrane. The upgrowth 

 of the outer supporting cells also forces axisward the apical 

 ends of the elements of the spiral organ and in this way con- 

 tributes a small part to the shift in the relative position of the 

 hair cells. A slight increase in width of the vestibular lip of 

 the spiral limbus may contribute a still smaller part by extending 

 the membrane outward." 



