PHANEROGAM I A 



495 



two scales are still more marked. The two anatropwis ovules are borne on au inner 

 scale, which, at its Lase, is continuous with the scale of the cone. THE OITKI: 



si .U.K. IS CAU.KH THE lUl.YCT SCALE, THE INNEK, THE OVULIFEKOUS SCALE (FigS. 



461, 462). The ovuliferous scale is the more strongly developed and it is the part 

 that becomes lignified and affords protection to the ovules. Even at the period 

 of flowering the bract scale is usually concealed by the ovuliferous scale and only 

 to be 'detected on close inspection. In other forms, however (e.g. Abies bracteata, 



Via. 4ii3. 1'M'a aci-ilx* (i nat. si/e). 1, Twix with male tloneis. ', Terminal female flown-, j, 

 Pendulous cone. !,, MicTosiM)iij]>liyll. .", MaCKMporophyll ; tlie bract-scale is covered by the 

 larjje, bent-back, ovuliferous scale ; an ovule is visible at the base of the ovuliferous scale. 

 '-', Kipe seed with the win;.: funned by a detacheil portion of the ovuliferous scale, (x 4-fi.) 



A. /ii'ctiiinta, Fig. 462 b, /'*< in/u/sni/ti limujJasi-i, etc.), the bract scales even in the 

 older cone project prominently between the ovuliferous scales. 



The view here followed, that the ovuliferous scale is a placental outgrowth of 

 the bract scale bearing the macrosporangia, and that its increase in size can be 

 traced in the series of the Araucarieae and Abietineae, is that supported by SACHS, 

 EICHLER, and GOEBEL. Attention must be drawn, however, to an alternative view 

 supported by STHASi;nif;Kii and CEUAKOV.SKY among others. According to this 

 the ovuliferous scale corresponds to the coherent bracteoles of an axillary shoot 

 borne in the axil of the bract scale. The cone itself would thus not correspond to 

 a single flower bearing a number of sporophylls, but to a system of shoots, i.r. it 

 would be an inflorescence. 



