Embryology. 129 



I will quote the following description of them, because, 

 for terseness combined with lucidity, it is unsur- 

 passable. 



Researches, chiefly due to Flemming, have shown that the 

 nucleus in very many tissues of higher plants and animals con- 

 sists of a capsule containing a plasma of" achromatin," not deeply 



FIG. 36. Karyokinesis of a typical tissue-cell (epifhelium of Sala- 

 mander). (After Flemming and Klein.) The series from A to I 

 represents the successive stages in the movement of the chromatin 

 fibres during division, excepting G, which represents the " nucleus- 

 spindle " of an egg-cell. A, resting nucleus ; D, wreath-form ; E, 

 single star, the loops of the wreath being broken ; F, separation of 

 the star into two groups of U-shaped fibres; H, diaster or double 

 star ; I, completion of the cell-division and formation of two resting 

 nuclei. In G the chromatin fibres are marked a, and correspond to 

 the " equatorial plate " ; b, achromatin fibres forming the nucleus- 

 spindle ; t, granules of the cell-protoplasm forming a " polar star." 

 Such a polar star is seen at each end of the nucleus-spindle, and is 

 not to be confused with the diaster H, the two ends of which are 

 composed of chromatin. 



stained by re-agents, ramifying in which is areticulum of "chro- 

 matin " consisting of fibres which readily take a deep stain. 

 ( Fig. 36, A). Further it is demonstrated that, when the cell is 

 about to divide into two, definite and very remarkable move- 

 ments take place in the nucleus, resulting in the disappearance 

 of the capsule and in the arrangement of its fibres first in the 

 * K 



