50 THEORY OF EVOLUTION 



The question as to whether a given character 

 is dominant or recessive is a matter of no theo- 

 retical importance for the principle of segre- 

 gation, although from the notoriety given to it 

 one might easily be misled into the erroneous 

 supposition that it was the discovery of this re- 

 lation that is Mendel's crowning achievement. 



Let me illustrate by an example in which the 

 hybrid standing between two types overlaps 

 them both. There are two mutant races in our 

 cultures of the fruit fly Drosophila that have 

 dark body color, one called sooty, another which 

 is even blacker, called ebony (fig. 20). Sooty 

 crossed to ebony gives offspring that are inter- 

 mediate in crolor. Some of them are so much 

 like sooty that they cannot be distinguished 

 from sooty. At the other extreme some of the 

 hybrids are as dark as the lightest of the ebony 

 flies. If these hybrids are inbred there is a con- 

 tinuous series of individuals, sooties, interme- 

 diates and ebonies. Which color here shall we 

 call the dominant? If the ebony, then in the 

 second generation we count three ebonies to 

 one sooty, putting the hybrids with the ebonies. 

 If the dominant is the sooty then we count three 



