DIVISION I. GENERAL MORPHOLOGY-. 



curved furrows ; and as gonidia are formed on the surface in a way that will be after- 

 wards described (see Division II) the body may be termed a gonidiophore (Fig. 16). 

 The hyphae of the Fungus-body must necessarily make their way for some distance 

 from the ovarian base into the floral pedicel, for it is difficult to conceive of any other 

 mode of supplying food to the Fungus ; but we have no exact information on this 

 point. When the gonidiophore is fully formed, the beginning of a sclerotium makes 

 its appearance in the torus at its base and on the mycelium, which is supposed to 

 spread through it, in the form of a small somewhat elongated fungal body enclosed 

 in the white tissue and distinguished by its greater density (Fig. 16 6, s). It is 

 formed at first of slender delicate separable hyphae which are continuous with 

 those in the vicinity, but are somewhat firmer and more closely compacted. Its 



FIG. 16. Claviceps ftirfurea, Tul. a young ovary of FIG. 17. Clavieeps furfiirta, Tul., on Secale certale. 



SetaU certale penetrated and covered with the gonidio- a seen from without, b median longitudinal section. The 



phore, seen from without; the hairs of the ovary and the sclerotium s rests on the torus and carries up the dry- 



remains of the style g project at the apex from the fungal ing gonidiophore f on its apex. After Tulasnc. Slightly 



investment, b longitudinal section through a similar stage magnified. 



in the development from Secale; s commencement of 

 the sclerotium. c similarly young state of the Fungus on 

 the pistil of Glyceria fluitans, the Fungus projecting 

 beyond the apex of the ovary. After Tulasne. Slightly 

 magnified. 



surface soon acquires a violet hue, the superficial cells beginning to assume the 

 character of the future rind. It now increases in thickness and elongates into the 

 well-known horn-shaped body, which is attached at its base to the torus and projects 

 above from between the paleae. The course of its development still requires more 

 exact investigation. Its growth in the longitudinal direction is no doubt maintained 

 by continued addition at the base. The increase in thickness of each transverse 

 section above the base mus^ in a great measure be due to the expansion of cells 

 already formed, since these are more than four times broader in the fully 

 developed parts than in the younger. The gonidiophore ceases to grow as soon as 

 the sclerotium begins to be formed, and being detached from the torus as the 

 sclerotium enlarges it is carried up like a cap on its apex, and there shrivels up and 

 sooner or later falls off (Fig. 1 7). 



