254 THE FLOWER. 



465. The connective may be appendaged either by a prolon- 

 gation or otherwise from the tip (as in Fig. -llt'J), or from the 

 back, as in Violets and in many Kricaceoiis plants. 



I (ill. The normal anther is two-celled, bilocular, or (to use 

 a less eoiiinion term) f/ifhecous, and its lohes or cells parallel, 

 right and left : but the cells at first, and sometimes at maturity, 

 are l>!l<i<-<'ll<itc. that i-> each is divided into two by a partition 

 which stretches from the connective to the suture or line of 



dehiscence. In an 

 innate anther, and 

 in main' others, this 

 line of dehiscence is 

 marginal or lateral, 

 either strictly or 

 nearly so, as in 

 Fig. 500. When introrse or extrorse (as in Fig. 5()1, 502), 

 the sutures may still be considered to represent the margins 

 turned inward or outward. The pollen is accordingly pro- 

 duced in four cavities or separate portions of the interior. But 

 *he two locelli on the same side of the midrib or connective 

 (right and left) are usually confluent into one pollen-tilled cavity 

 or cell at maturity if not earlier, or at least the partition between 

 hem breaks up at dehiscence. Sometimes it remains, and, the 

 groove at the sutures being deep, the anther is strongly four- 

 lobed or quadrilocular at maturity, as in Menispermum (Fig. 

 504) ; but morphologically this is still only bilocular (dithecous) 

 although <|uadrilocellate, and the anther opens at the sutures 

 and through these partitions. 



467. A stamen being the homologue of a leaf, the natural 

 supposition is that the anther is homologous with the blade or 

 an apical portion of the blade, therefore the two lobes or thcca- 

 with the right and left halves of it. the intervening connective 

 with the midrib, and the line of dehiscence with the leaf- mar- 

 gins. 1 This conception is exemplified by the accompanying 



1 'Phis is the view lonir airo taken by Cassini and Ku'per. and it may still 

 lie maintained a- the best morphological conception. Mohl interposed some 



dltjertiniis in its universality: luit, as presented in Sadie's Text-Hook, they 

 are not incompatible with tin 1 common morphology. Sachs takes the fi la- 

 ment with the connective to be the liomoloL'ue of the whole leaf, and the 

 anther-cells as appendages. Others, in likening the anthers to glands, adopt 

 a similar vie \v. 



FIC. .".on. I mint i- aiitlicr. smiie us Kit,'. l!H - .. in \ ounu'rr -t:iti-. \\illi transverse section, 

 sliowiii'j tlir f'Hir li'ivlli. r.ni. Same of an adn.'ite extrorso antlir-r. snc-li as Fig. 407. 

 ">()?. S:inir :is the priM-ciliiiu' luit mature ainl ilehisccut, the two locelli becoming one cell 

 l>y the vanishing <>r lireaking up of the partition. 



