CAPRIFOLIACE^E. 



ORDER LXIX. CAPRIFOLIACE^E. 



Shrubby, or a few perennial herbaceous plants, with opposite leaves normally 

 destitute of stipules, and regular or (in the corolla) irregular hermaphrodite flow- 

 ers ; calyx-tube adnate to the 2-5-celled or by suppression 1-celled ovary ; sta- 

 mens as many as lobes of the corolla (in Linncea one fewer, in Adoxa doubled) 

 and alternate with them, inserted on its tube or base ; embryo small in the axis 

 of fleshy albumen. Corolla-lobes generally imbricated in the bud. Ovules ariatro- 

 pous, when solitary suspended and resupinate ; the rhaphe dorsal. Seed-coat 

 adherent to the albumen. Flowers commonly 5-merous. 



TRIBE I. SAMBUCE^E. Corolla regular, short, rotate or open-campanulate, 5-lobed. 

 Style short or hardly any : stigmas 3 to 5. Ovules solitary in the (1 to 5) cells. Fruit 

 baccate-drupaceous ; the seed-like nutlets 1 to 5. Inflorescence terminal and cymose. 



* Herb, with stamens doubled and flowers in a capitate cluster. Anomalous in the order. 



1. ADOXA. Calyx with hemispherical tube adnate to above the middle of the ovary; limb 

 about 3-toothed. Corolla rotate, 4-6-cleft. Stamens a pair below each sinus of the corolla, 

 each with a peltate one-celled anther, and the short subulate filaments approximate or united 

 at base (one stamen divided into two). Ovary 3-5-celled : style short, 3-5-parted. Ovule 

 suspended from the summit of each cell. Fruit greenish, maturing 2 to 5 cartilaginous nut- 

 lets. Cauline leaves a single pair; radical ones and scales of the rootstock alternate ! 



* * Frutescent to arborescent : inflorescence compound-cymose : flowers articulated with 

 their pedicels : stamens as many as corolla-lobes : anthers 2-celled : calyx 5-toothed. 



2. SAMBUCUS. Leaves pinnately compound. Corolla rotate or nearly so. Ovary 3-5- 

 celled, forming small baccate drupes with as many cartilaginous nutlets. Embryo nearly 

 the length of the albumen. 



3. VIBURNUM. Leaves simple, sometimes lobed. Corolla rotate or open-campanulate. 

 Ovary 1-celled and 1-ovuled, becoming a drupe with a single more or less flattened nutlet or 

 stone. Embryo minute. Cymes in some species radiate. 



TRIBE II. LONICERE^E. Corolla elongated or at least campanulate, commonly more 

 or less irregular. Style elongated : stigma mostly capitate. Fruit various. Stipules 

 or stipular appendages seldom seen. 



* Herbs, with axillary sessile flowers and drupaceous fruit. 



4. TRIOSTEUM. Calyx-lobes 5. Corolla tubular-campaimlate, somewhat unequally 5- 

 lobed ; tube gibbous at base. Stamens 5. Ovary 3- (sometimes 4-5-) celled, with a single 

 suspended oviile in each cell : style slender : stigma 3-lobed. Fruit a fleshy drupe, crowned 

 with the persistent calyx-lobes : putamen bony, costate, at length separable into 3 (rarely 4 

 or 5, or by abortion 2) thick one-seeded nutlets. 



* * Fruticulose creeping herb, with long-pedunculate geminate flowers and dry one-seeded 

 fruit, but a 3-celled ovary. 



5. LINN-3EA. Calyx with limb 5-parted into subulate-lanceolate lobes, constricted above the 

 globular tube, deciduous from the fruit. Corolla campanulate-funnelform, not gibbous, al- 

 most equally 5-lobed. Stamens 4, two long and two shorter, included. Ovary 3-celled; two 

 of the cells containing several abortive ovules ; one with a solitary suspended ovule, forming 

 the single seed in the dry and indehiscent coriaceous 3-celled small fruit. Style exserted : 

 stigma capitate. 



* * * Shrubs, with scaly winter-buds, erect or climbing : fruit 2-many-seeded : style slen-. 

 der : stigma capitate, often 2-lobed. 



6. SYMPHORICARPOS. Calyx with a globular tube and 4-5-toothed persistent limb. 

 Corolla regular, not gibbous, from short-campanulate to salverform, 4-5-lobed. Stamens as 



