THE ALIMENTARY CANAL 



357 



less removed from the lumen they possess an extensive system of peri- 

 cellular secretory canals which invest the cell in a basket-like manner 

 and convey its secretion to the glandular lumen, where it mixes with the 



secretion of the chief cells. The parietal cells 

 also possess a system of intracellular canaliciili. 

 The parietal cells are commonly believed 

 to secrete the HC1 of the g:isiric juice. But 

 Harvey and Bensloy (Biol. Bull., 23, 4, 1912) 

 claim to have shown 

 that free HC1 is not 

 present in these cells; 

 their content is said to 

 be chemically neutral 

 or alkaline, and to con- 

 sist largely of chlorids. 

 The results of experi- 

 ments with rabbits and 

 various other verte- 

 brates indicate that 

 chloriu is secreted by 

 the parietal cells in the 

 form of a chlorid of an 

 organic base, and that 

 the HC1 is only set free 

 after the secretion is 

 poured out of the gland 

 into the foveola. Ham- 

 mett (Anat. Rec., 9, 1, 

 1915), however, presents 

 further evidence tend- 

 ing to show the pres- 

 ence of acid in the pari- 

 etal cells. 



PYLORIC GLANDS. 

 These are branched con- 

 voluted tubular glands with relatively long crypt-like ducts, into the 

 bottom of which several secreting tubules open. According to Piersol, 

 they occupy the pyloric fifth of the stomach. 



The typical convolution is found only in the fundus of the gland, 

 the course of the ducts being nearly straight. The branching, on the 



FIG. 333. A PYLORIC 

 GLAND, FROM SECTION 

 OF THE DOG'S STOMACH. 

 (Ebstein.) 



m, mouth; n, neck; tr, a 

 deep portion of a tubule 

 cut transversely. (From 

 "Quain's Anatomy.") 



FIG. 334. PORTION OF 

 GASTRIC GLAND 

 FROM THE FUNDUS 

 REGION OF THE 

 STOMACH. 



L, lumen, ending in 

 intracellular secretory 

 canaliculi in the parie- 

 tal cells (P.c.); C.c., 

 chief cells. Prepared 

 by the Golgi chromate 

 of silver impregnation 

 method; highly magni- 

 fied. (After Zimmer- 

 man n.) 



