158 ANTHOCYANINS AND GENETICS 



Bateson (480) suggested that a better interpretation of de Vries' 

 results would be to suppose that the Fj plants produced, in each sex, 

 in equal numbers, gametes having the characters, R, F, D and W, 

 and these, on fortuitous mating, would give 9R : 3F : 3D : 1W. In 

 view of more recent work, both views have to be revised. 



In 1907 the author (514) published results of work on a number of 

 colour- varieties ; those used were 



1. White lips and tube of the corolla pure white. 



2. Yellow lips yellow; tube ivory. 



3. Ivory -lips and tube ivory. 



4. Crimson lips crimson; tube magenta. 



5. Magenta lips and tube magenta. 



With the exception of white, all varieties have a constant orange- 

 yellow palate. Further varieties occur in which the lips are magenta 

 or crimson, and the tube is ivory; the term 'delila' was used by de 

 Vries for these varieties and has been retained. 



The inheritance of colour in the varieties employed was represented 

 by the following factors : 



Y. A factor representing yellow colour in the lips associated with 

 ivory tube-colour. 



I. A factor representing ivory colour in the lips. 



L. A factor representing magenta colour in the lips. 



T. A factor representing magenta colour in the tube. 



It was found also that certain relationships existed between the 

 factors, i.e. 



1. All zygotes, from which Y is absent, are white, though they 

 may contain any of the factors I, L and T. 



2. The effects of the factor T are not manifested unless L is also 

 present in the zygote ; that is, no magenta colour appears in the tube 

 unless there is magenta colour in the lips. 



3. All zygotes containing Y are coloured, the actual colour being 

 modified and determined by the presence of one or more of the remaining 

 factors. A zygote containing Y only, or Y and T is yellow. 



4. Ivory is dominant to yellow; a zygote containing Y and I, or 

 Y, I and T is ivory. 



5. Since magenta superposed upon yellow gives crimson, a zygote 

 containing Y, L and T is crimson, Y and L only, crimson delila. 



6. Magenta upon ivory gives magenta. A zygote containing 

 Y, I, L and T is magenta, Y, I and L only, magenta delila. 



A further variety termed ' rose ' was employed, but full details of 



