ANTHOCYANINS AND GENETICS 189 



nearly complete deficiency up to such predominancy as almost to hide 

 the pale yellow ground-color " (de Vries, 498). 



Papaver orientale. The dark patches of anthocyanin at the base 

 of the petals are absent in some varieties (de Vries, 565). 



P. Rhoeas. In some varieties there is a white margin to the petals 

 due to the presence of a dominant inhibiting factor (Shull, 588). 



P. somniferum. Some varieties have dark basal patches of pigment 

 (anthocyanin) on the petals. From others it is absent. On crossing 

 dark "'Mephisto' with the white-hearted 'Danebrog' the hybrid 

 shows the dark pattern" (de Vries, 498). 



Phaseolus. In some strains the mottling of anthocyanin in the 

 testa is due to a factor M, but the pattern only shows when the plant 

 is heterozygous for M. In other strains the mottling occurs in both 

 heterozygous and homozygous forms (Emerson, 528, 529; Shull, 513, 

 521 ; Tschermak, 479, 590). 



Pisum. The mottling of anthocyanin in the testa is due to a definite 

 Mendelian factor (Lock, 518). 



Primula sinensis. The patches of anthocyanin outside the yellow 

 eye are a definite Mendelian character. They do not develop to their 

 fullest intensity unless the stigma is red, or the flower is deep-coloured ; 

 the spots are pale in pale-coloured flowers and in flowers with a green 

 stigma. 



The 'Duchess' distribution of pigment, i.e. a ring of colour round 

 the eye, spreading to a flush, may be regarded as pattern. So also 

 the 'Sirdar' variety, where the colour is present in minute dots giving 

 the flower a dusty appearance, the margins of the petals being white 

 (Gregory, 557). 



Solanum tiiberosum. The flowers of several white varieties have 

 'tongues' of colour radiating out from the throat to the tips of the 

 corolla segments. The 'tongues' show different degrees of intensity 

 of coloration, governed, probably, by different factors (Salamau, 

 unpublished work). 



Tropaeolum majus. There is at present no published work on the 

 flower colour of this genus but, as far as observations go, there is a 

 definite inheritance of the blotch at the base of the petals. The original 

 type has an orange-red (anthocyanin and yellow plastids) flower, rather 

 deeper in colour at the base of the petals, and with dark ' honey-guides ' 

 running into the 'claw.' There are varieties in which the anthocyanin 

 has disappeared from the main part of the flower so that the flower 

 is yellow with orange-red blotches at the base of the petals. If the 



