194 ANTHOCYANINS AND GENETICS 



95-98 %, and they give rise to the red type either by seed or bud 

 variation. 



4. The broadly-striped individuals produce more reds than the 

 narrow-striped, the average being about 39 %. 



5. The red individuals thrown by the striped resemble the red 

 type, but differ in again throwing striped. The intensity of inheritance 

 of the red character is about 70-85 %. 



6. The yellow variety (without anthocyanin) does not arise from 

 the striped race. 



De Vries draws attention to the fact that striped varieties do not 

 occur in nature. Other examples he gives of striping are in Stocks, 

 Liver-leaf (Hepatica), Dame's Violet (Hesperis}, Sweet William (Dianthus 

 barbcttus) and Periwinkle ( Vinca minor) ; also in Hyacinthus, Cyclamen, 

 Azalea, Camellia, and in the Meadow Crane's-bill (Geranium pratense) 

 when cultivated. Also these, he says, are known to come true to 

 striping when seed is taken from striped individuals, and from time 

 to time to throw self-coloured individuals. He made pedigree cultures 

 of the Dame's Violet (Hesperis matronalis) for five years, and of Clarkia 

 pulchella for four years, and they both behaved in exactly the same way 

 as Antirrhinum. He further points out that other parts of plants may 

 be striped, as, for instance, the red and white striped roots of Radishes, 

 and the inflorescence of the Cockscomb (Celosia cristata). 



From these results one would gather that. the inheritance of striping 

 is non-Mendelian. More recently Emerson (605) has suggested that 

 striping, on the basis of a certain hypothesis, may conform to the 

 Mendelian scheme of inheritance. It appears that ears of certain 

 varieties of Maize show striping in the red pigment developed in the 

 pericarp. Emerson says : " Plants in which this pigment has a variegated 

 pattern may show any amount of red pericarp, including wholly self- 

 red ears, large or small patches of self-red grains, scattered self-red 

 grains, grains with a single stripe of red covering from perhaps nine- 

 tenths to one-tenth of the surface, grains with several prominent stripes 

 and those with a single minute streak, ears with most of the grains 

 prominently striped and ears that are non-colored except for a single 

 partly colored grain, and probably also plants with wholly self-red 

 and others with wholly colorless ears." A number of selfmgs were 

 made for several generations of both homozygous and heterozygous 

 variegated cobs ; homozygous and heterozygous variegated cobs were 

 also crossed with true-breeding white male plants. From his experi- 

 ments Emerson deduces the following results: 



