ANTHOCYANINS AND GENETICS 161 



The different varieties may be represented as: 



yyI(i)I(i)L(l)L(l)T(t)T(t)D(d)D(d)B(b)B(b) white. 

 YY(y)iillT(t)T(t)D(d)D(d)B(b)B(b) yellow. 

 YY(y)II(i)llT(t)T(t)D(d)D(d)B(b)B(b) ivory. 

 YY(y)iiLL(l)ttddbb yellow tinged bronze delila. 

 YY(y)iiLL(l)TT(t)ddbb yellow tinged bronze. 

 YY(y)II(i)LL(l)ttddbb -ivory tinged rose dore delila. 

 YY(y)II(i)LL(l)TT(t)ddbb ivory tinged rose dore. 

 YY(y)iiLL(l)ttDD(d)bb bronze delila, 

 YY(y)iiLL(l)TT(t)DD(d)bb-bronze. 

 YY(y)II(i)LL(l)ttDD(d)bb rose dore delila. 

 YY(y)II(i)LL(l)TT(t)DD(d)bb rose dore. 

 YY(y)iiLL(l)ttDD(d)BB(b) crimson delila. 

 YY(y)iiLL(l)TT(t)DD(d)BB(b) crimson. 

 YY(y)II(i)LL(l)ttDD(d)BB(b) magenta delila. 

 YY(y)II(i)LL(l)TT(t)DD(d)BB(b) magenta, 



Certain varieties deeper .than intermediate magenta are known 

 to exist; they are recessive to intermediate magenta, but otherwise 

 they have been very little worked with. The heterozygous forms 

 have already been mentioned above. The original wild type is 

 thought to be intermediate magenta and would have the constitution 

 YYIILLTTDDBB. As would be gathered from the above statements, 

 ivory x white will give magenta ; hence two albinos, as regards antho- 

 cyanin, will give colour. Whether or no this case is strictly comparable 

 to the results of crossing two whites in Lathyrus and Matthiola will be 

 discussed later. 



Arum maculatum. A. maculatum occurs in nature in two forms; 

 one has black (anthocyanin) spots on the leaves, the other is without 

 spots. Colgan (552) obtained seeds from a spotted plant, and found, 

 out of 11 seedlings, 5 were spotted and 6 unspotted. Colgan suggests 

 the female parent was heterozygous for spotted character and was 

 fertilised by the unspotted form, but there was no evidence in support of 

 the suggestion beyond the fact of equality of forms among the offspring. 



Aster Tripolium. De Vries (498) notes that pigmented type is 

 dominant to albino. 



Atropa Belladonna. Saunders (475) made crosses between A. Bella- 

 donna typica and A. Belladonna var. lute.a. The type has brown 

 (anthocyanin on plastid) flowers, black (anthocyanin) fruits, and stem 

 tinged with anthocyanin: the variety, which is without anthocyanin, 



w. p 11 



