ANTHOCYANINS AND GENETICS 181 



Beta. Some varieties have leaves coloured quite red with antho- 

 cyanin. This only happens in red-fleshed varieties, i.e. when antho- 

 cyanin is formed in the inner tissues, as well as in the skin, of the root. 

 In varieties having colourless flesh, the leaves are green (Kajanus, 559). 



Corchorus capsularis. Varieties with deep red (anthocyanin) stems, 

 petioles and fruits have the teeth of the leaves also tipped with red. 

 Other strains with less pigment in the stem have no colour in the leaf 

 teeth (Finlow & Burkill, 572). 



Datura. D. Tatula has violet flowers and red stems (anthocyanin). 

 D. Stramonium has white flowers and green stems (Saunders, 475, 487). 



Digitalis purpurea. The presence of a factor, M, for magenta pig- 

 ment always brings with it colour in the spots on the lower inner surface 

 of the corolla. Even in the presence of M the general colour in the 

 corolla may be inhibited by another factor I, but the spots remain 

 red (var. white spotted with magenta). If M is absent, the spots are 

 colourless (Keeble, Pellew & Jones, 542; Saunders, 563). 



Gossypium. In the Indian cotton, strains having yellow flowers, 

 or white flowers with only a basal spot on the petals, have green foliage. 

 In strains having red pigment, the varieties having red flowers (forms 

 homozygous for the reddening factor, see p. 166) have the veins and 

 lamina of their leaves suffused with red. In leaves of the plants which 

 have flowers red on yellow, or red on white (forms heterozygous for the 

 reddening factor) the anthocyanin is confined to the veins (Leake, 561). 



Helianthus. In the wild H. annuus the disk is purple, the colour 

 (anthocyanin) being in the tips of the paleae, the margin of the corolla 

 and the styles and stigmas. The 'Russian Sunflower' has the tips of 

 the paleae yellowish-green, the corolla, styles and stigmas yellow 

 (Shull, 520). 



The chestnut-red-flowered Sunflower, which has anthocyanin on 

 yellow in the ray florets, has dark reddish-purple stems. The purple- 

 flowered variety, anthocyanin on primrose, has no reddish-purple in 

 the stems (Cockerell, 611). 



Lathyrus odoratus. The albinos always have green leaf-axils. 

 When a certain factor for production of colour in the axils is present 

 and the flowers are coloured, there is a reddish-purple spot in the axil ; 

 when the axil colour factor is absent, the axils are green (Bateson, 524). 

 Red tendrils are (?) always associated with red in the axils of leaves 

 (Bateson, 499). 



Oenothera. In the variant Oenothera rubricalyx red buds and 

 hypanthia are said to be associated with red stems (Gates, 555). In 



