182 ANTHOCYANINS AND GENETICS 



the Fj, however, from reciprocal crosses between 0. rubricalyx x 0. 

 Lamarckiana, plants appeared in which red buds were associated 

 w r ith a low degree of red pigmentation in stems and rosettes, whereas 

 pale red buds and green hypanthia were associated with brilliant red 

 stems, and buds entirely free from anthocyanin with dark stems 

 (Shull, 609). 



Pisum sativum. White-flowered plants have no colour in the leaf- 

 axils and no red or purple (anthocyanin) colour in the testas. Red- 

 flowered plants have red colour in the leaf-axils and red spots on the 

 testa. Purple-flowered plants have purple colour in the leaf-axils 

 and purple spots on the testa (Lock, 518). 



Primula sinensis. Recessive white-flowered varieties may have 

 green or red stems (Keeble & Pellew, 541). Pale-flowered varieties 

 may be on green or faintly coloured stems; deep-flowered varieties 

 appear on deep red stems only; whereas light-flowered varieties may 

 be borne on either deep- or light-stemmed plants. The 'Sirdar' variety, 

 in which the pigment is in minute dots and the edges of the petal are 

 white, has stems with a red 'collar,' i.e. base of stem. If the flower- 

 colour is crimson or magenta, the stem colour is purplish-red ; if the 

 flower is true red ('Orange King') the stem pigment is also true red. 

 Blue flowers have blue stem pigment. 



The deep spots of colour external to the eye are not fully developed 

 unless the stigma is coloured. They are deeply-coloured only in 

 deeply-coloured flowers : in light flowers they are similar to those in 

 flowers with a green stigma. They also depend on base colour as they 

 are not present in pale flowers, nor in striped flowers, unless the stripe 

 occurs in the area occupied by the spot (Gregory, 557). 



Salvia Horminum. The bracts of the inflorescence are purple, 

 red, or green according to whether the flowers are purple, red, or white 

 (Saunders, 487). 



Zea Mays. Red pigment may appear in the pericarp, cobs and silks 

 (stigmas) separately and independently in each part. If the glumes 

 are red, red is present in some other part of the plant, though it may 

 only be in the silks (East & Hayes, 553; Emerson, 554). 



The various relationships between flower-colour and the presence 

 of anthocyanin in the vegetative organs may therefore be classified 

 as follows: 



1. Simple case. Loss of colour in the flower accompanied by total 

 loss from vegetative organs. Ex. : true albinos of Antirrhinum, Atropa, 

 Lathynts, Pisum, Primula, etc. 



