ANTHOMEDUS.E PROBOSC1 DACTVI.A. ISO 



The type species of this genus is Proboscidactyla flavifirrata Brandt, from the North 

 Pacific. Its 4 mam radial-canals branch more profusely than in any other species of the genus. 



The American species, P. ornntn, was first described by McCrady under the name H'illsia 

 ornata from Charleston Harbor, South Carolina. 



GENERIC CHARACTERS. 



Proboscidactyla: Dendrostaurinae having 4 primary radial-canals which give rise to simple, 

 or branched, side branches during the growth of the medusa. The tentacles are simple and 

 hollow and are situated at the extremities of the radial-canals and their branches. Upon the 

 exumbrella there are clusters of nematocysts which alternate in position with the tentacles. 

 There are no marginal clubs or cirri and there is no circular canal. The gonads are upon 

 the 8 adradial sides of the stomach extending outward along the sides of the 4 radial-canals. 



Haeckel, 1879, separates that which we here designate as Proboscidactyla into 4 genera 

 as follows: 



l. Dicranocanna, in which the 4 radial-canals branch once, giving 8 radial vessels which reach the bell-margin. 



2. Dyscannota, in which the 4 primary radial-canals give rise each to 1 side branches, and thus ll canals reach the 



bell-margin. 



3. H'lllfila, in which the 4 original radial-canals give rise each to 2 side branches, and the first-formed side branch uf 



each quadrant itself gives rise to a side branch, so that 1 6 canals reach the bell-margin. 



4. I'rohoscidactyla, in which the 4 canals that rise from the stomach branch complexly, and branches arise from the pri- 



mary branches, etc., so that 32 or more radiating canals reach the bell-margin. 



It is evident that medusae of the genus ProbosciJactyla may successively pass through 

 stages represented by the genera Dicranocanna, Dyscannota, and H'illettn. 



The American, Atlantic Proboscidactyla or nut a is sexually mature in the " H'llletta" stage, 

 but its southern variety gemmifera gives rise to medusa-buds while in the "D\scannota" stage. 



Psythia prolijera, Agassiz and Mayer, 1902, Mem. Museum Comp. Zool. at Harvard 

 College, vol. 26, p. 143, plate i, fig. 3, may be a young budding Proboscidactyla? 



Proboscidactyla flavicirrata Brandt. 



Proboscidactyla flaricirrata, BRANDT, 1854, Recueil Actes seances publiques Acad. Imp. Sci. St. Petersbourg, p. 28 (of the "Separ- 

 ate"); 1838, Mem. Acad. Imp. Sci. St. Petersbourg, Sci. Nat., ser. 6, tome 4, Par. 2, p. 390, plate 19. AIIASSIZ, A., 1865, 

 North Amer. Acal., p. 173, figs. 280-282. -BROWNI:, 1904, Fauna and Geography Maldive and I.accadive Archipelagoes, 

 vol. 2, p. 725. BKDOT, 1905, Revue Suisse de Zool., tome 13, p. 144 (all papers to 1850). 



ProboiciJactyla fiaricirrata-^- P. brfvictrrata, HAECKEL, 1879, Syst. der Medusen, pp. 159, 160. 



ProbotciJactyta brti'icirrata, MIIRHACH and SHEARER, 1903, Proc. Zool. Soc. London, vol. 2, p. 178. 



Bell thick-walled and dome-like to globular. About 12 mm. wide and 10 mm. high. 

 4 main radial-canals arise from the stomach, but these fork near their points of origin and 

 each branch gives rise to side branches, from one side only and these again give off similarly 

 arranged branches so that about 54 to 70+ terminal branches reach the bell-margin. There 

 is a short tentacle at the end of each terminal branch of the radial-canals. There are clusters 

 of nematocysts on the exumbrella alternating with the tentacles. There is no ring-canal. 

 Stomach small, with 4 recurved lips. Cionads upon the sides of the 4 radial pouches of the 

 stomach. There are neither medusa-buds nor stolons. 



Stomach, gonads, and tentacle-bulbs dull yellow. North Pacific, coast of Kamtschatka 

 and Puget Sound, Washington. 



This species is distinguished by the profuse and peculiar branching of its radial-canals. 



Proboscidactyla ornata Browne. 

 Plate 20, figs. 1-10. 



H'illiia ornata, McCRADY, 1857, Gymn. Charleston Harbor, p. 47, plate 9, figs. 9-11. 



H'illia ornata, AGASSIZ, L., 1862, Cont. Nat. Hist. U. V, vol. 4, p. 34(1. AGASSIZ, A., 1862, Proc. Boston Soc. Nat. Hist., vol. 

 9, p. 96, figs. 20, 21 ; 1865, North Amer. Acal., p. 171, figs. 2740-279. VERRILL, 1873, Report Commiss. Fish and Fisheries 

 U. S. for 1871-72, pp. 455, 735. FEWCU, 1882, Bull. Mus. Cump. /mil. at HarvaYd College, vol. 9, p. 299, figs. 22, 23, 

 non fig. 24; 1884, Mem. Mus. Comp. Zool. at Harvard College, vol. 9, No. 3, plate 5, fig. 19. MAVF.R, 1904, Mem. 

 Nat. Sci. Museum Brooklyn Inst. Arts and Sci., vol. l, No. l, p. 13, plate 2, fig. 12. NUTTING, 1901, Bull. U. S. Fish 

 Commission for 1899, vol. 19, p. 377. HARGITT, 1904, Bull. U. S. Bureau of Fisheries, vol. 24, p. 40, plate i, fig. 5. 



Dvscannota JvsJipleura, HAECLEL, 1879, Syst. der Medusen, p. 152. 



H'illetta ornata, HAECKEL, 1879, Syst. der Medusen, p. 157. 



ProboscvJactyla ornata, BROWNE, 1904, Hydromedusa? Maldive Islands, p. 726.- MAAS, 1904, Sitzungsber. math.-phys. Klasse 

 kgl. Bayer. Akad. Wissenschaft., Bd. 34, p. 439. 



( ?) Dicranocanna furcitlata, HAECKEL, 1879, Syst. tier Medusen, p. 156. 



