CORONAT/E PARAPHYLLINA. 549 



Genus PARAPHYLLINA Maas, 1903. 



Parapln'llina, MAAS, 1903, Scyphomedusen der Siboga Expedition, Monog. 1 1, p. 6; 1907, Ergeb.urul Fortschritte dcr Zool., Bd. I, 

 p. 195. 



The type species is Paraphyllina intermedia Maas, 1903; from the Malay Archipelago, 

 between 100 fathoms and the surface. 



GENERIC CHARACTERS. 



Coronatae with 12 tentacles, 4 perradial marginal sense-organs, and 1 6 lappets. The 12 

 tentacles are interradial and adradial with reference to the stomach and lips. There are 

 4 pairs (8) of interradial gonads. The marginal sense-organs have a terminal lithocyst-sac 

 and a ventral bulb-like swelling, just beyond which is an eye with ectodetmal lens and ecto- 

 dermal pigment. 



The short 4-sided throat-tube and flat disk-like bell resemble the condition noted in 

 Nausithoe. but in its 4 marginal sense-onrans and 12 tentacles it recalls the condition seen in 



i O O 



Periphylla, with the important difference that in Periphylla the sense-organs are interradial, 

 whereas in Paraphyllina they are perradial in position. 



Maas places this genus in a new family which he calls the Paraphyllinidae. The characters 

 of this family are those ot its only genus, Paraphyllina. It is closely related to the fossil genus 

 Paraphyllites, Maas, 1906 (Neuen Jahrbiich. tiir Mmeralogie, Geol. und Palaontol., Bd. 12, 

 p. 90, 4 fign.). This lossil medusa differs from recent Paraphyllina only in that its 8 gonads are 

 adradial and placed 45 apart, whereas in Paraphyllina they are grouped in pairs on both 

 sides of the 4 interradii. 



Paraphflhtrs disttnetus is described by Maas from a well-preserved specimen. It is 

 15 mm. wide and has a well-developed coronal furrow, 16 pedaha, 4 perradial marginal 

 sense-organs, 12 tentacles, and 8 adradial gonads 45 apart. It is from the lithographic slate 

 of Kelheim. The latest reference to this fossil is that of von Ammon, 1908, Geonostischen 

 Jahiespeften, Jahrg. 19, p. 170. 



Paraphyllina intermedia Maas. 



Paraphyllina intermedia, MAAS, 1903, Scyphomeduscn dcr Siboga Expedition, Monog. II, p. 8, taf . 2, fign. 10-14; taf. 1 1, fig. 106. 

 Periphylla dodecabostrycha, LOBIANCO, 1903, Mitth. Zool. Sta. Neapel, Bd. 16, p. 219, taf. 7, fig. 3. See aku MAAS, 1^04, Result. 

 Camp. Sci. Prince de Monaco, fasc. 28, p. 48, foot-note. 



Bell flatly rounded, 8 mm. high and 15 mm. wide, without pointed apex. The coronal 

 furrow around the exumbrella is deep and separates the smooth, central, flatly-rounded apex 

 of exumbrella from the marginal zone ot lappets. Lappet-zone about as wide as radius of 

 central disk of exumbrella. The 16 pedaha ot the marginal zone are rectangular with rounded 

 angles and are separated one from another by deep radiating furrows. These pedalia are 

 in the radii of the tentacles and sense-organs and alternate with the lappets. The 12 pedalia 

 of the tentacles are of uniform width, while the 4 pedalia of the marginal sense-organs are 

 only about half as wide as the former. The 16 marginal lappets are oval and bluntly pointed, 

 and the 8 lappets flanking the 4 sense-organs are somewhat narrower than the others. The 

 12 tapering, hollow tentacles are all of equal length and are not quite as long as the diameter 

 of the bell. 



The 4 marginal sense-organs are each covered by a hood-like fold of the exumbrella. 

 Each sense-club contains a small terminal sac-shaped entodermal concretion. On the ventral 

 (subumbrella) side of the sense-club is a large ectodermal eye with a cup-like mass of pig- 

 ment and a spherical lens. On the inner side of the eye and upon the ventral side of the 

 sense-club is a bulb-like swelling. Altogether the sense-clubs resemble those of Nausithoe, 

 but the eye is larger. 



The throat-tube is short and 4-sided and the mouth is a simple cruciform opening. There 

 are 4 interradial rows of gastric filaments. The coronal ring-canal gives rise to 16 peripheral 

 pouches in the radii of the sense-organs and tentacles, and these are put into communication 

 one with another by means of a marginal ring sinus, as in Nausithoe. 



The gonads resemble those of Palephyra and are intermediate in character between 

 those of Periphylla and those of Nausithoe. They consist of 4 pairs of bean-shaped or egg- 

 shaped sacs. These 8 sacs project from the subumbrella floor on both sides of the 4 inter- 

 radii near the sides of the throat-tube and centripetal to the ring-muscle. 



