THE ORIGIN AND DEVELOPMENT OF GASTRIC GLANDS. 



6 9 



subject. The oesophageal glands of Urodcla have received more 

 or less consideration as the question of the homology of these 

 glands, with one another, and with those of higher forms has for 

 a long time aroused much interest. But, leaving this interesting 

 problem entirely out of the discussion, there are described in the 

 stomach of some of the Urodela two kinds of glands, the an- 

 terior and posterior oxyntic glands of Langley ('Si). Bensley 

 ('oo), for Amblystoina, and Carlier ('98), for Triton, have given 

 excellent descriptions of these glands. In the adult form the 



2 



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jSjjMjiir''-"' 



FIG. I. Transection through the fundus of the stomach of a larval Dcsmogna- 

 tJnis, 11.5 mm. Fixed in Gilson's, modified, a., gland anlage, "ovoid cell," just 

 about to divide; b.m., basement membrane; d., insinking at the surface of the 

 epithelium to form the future excretory duct ; ep., epithelial lining of the stomach, 

 showing one of the cells in process of division; ;;/., mesodermic tissue forming the 

 muscular layers about the stomach. This also shows mitoses. Obj. y'g, Oc. I. 



glands consist of a duct, composed of cells resembling the sur- 

 face epithelium, a neck, composed of clear mucous cells, and a 

 body, made up of granular cells containing large nuclei. The 

 glands of the cardiac, fundic and pyloric regions vary in complex- 

 ity but all resemble, to a greater or less extent, the type form 

 just described. These anterior oxyntic, or cardiac glands, may or 

 may not be transition forms between the true gastric glands and 



