FLAGELLATE FAUNA OF CCECUM OF SQUIRREL. 289 



the body. Measurements indicate a diameter of from 3 to 4.5 

 micra. Anterior to the nucleus lie a number of deeply staining 

 granules, concerning the affinities of which there is so much 

 controversy. Kofoid and Swezy's account of the arrangement 

 of these granules based largely upon their observations of the 

 cyst is somewhat as follows. There lies upon the nuclear mem- 

 brane a centrosome, which is connected by the "nuclear rhizo- 

 plast" with a "primary blepharoplast." This blepharoplast is 

 joined to the "secondary blepharoplast" by means of the 

 "transverse rhizoplast.". Finally, another thin thread-like line 

 connects the secondary and tertiary blepharoplast. Directed 

 somewhat posteriorly from the secondary and tertiary rhizoplast 

 are two deeply staining fibrils, the one (parastyle) supporting the 

 left lip of the cytostome, the other (parabasal body) supporting 

 the right lip. Two of the three anterior flagella take their origin 

 in the primary blepharoplast, one from the secondary. From the 

 tertiary rhizoplast there arises, in addition to the "parabasal 

 body," the cytostomal flagellum and a thread enclosing the 

 mouth, the "peristomal fiber." The cytostomal flagellum 

 vibrates along the edge of an undulating membrane. Dobell 

 and O'Connor deny the existence of any integrated fibrillar 

 system (neuromotor system), or connections between the granules 

 at the anterior end, claiming in addition that there are six 

 granules instead of four. Furthermore, in regard to the peri- 

 stomal fiber they state, "We believe there is no such fibre, and 

 that the mouth is not situated in this position." 



Despite long and painstaking study the writer was not able to 

 make out all the details of the granule complex with the positive- 

 ness of Kofoid and Swezy. That there is an integrated fibrillar 

 system connecting the granules, however, the writer has no doubt. 

 The rhizoplast connecting the blepharoplasts designated by 

 Kofoid as secondary and tertiary is especially prominent in most 

 specimens because of its thickness and deeply-staining qualities 

 (Fig. 12). A fine rhizoplast running from a granule on the 

 nuclear membrane was occasionally recognized. The best evi- 

 dence for the fibrillar system, however, comes from the dividing 

 stages, which will be described below. The writer was not able 

 to make out the exact affinities of the flagella, because of the 



