STUDIES IN THE LIFE-HISTORY OF EUGLENA. 333 



interesting information can be had regarding the development of 

 the motor or kinetic complex. Each nucleus is seen to be some- 

 what pear shaped, its apex turned toward the border of the new 

 reservoir and usually in contact with it (Fig. 13). At the apex 

 of each nucleus there appears the deeply staining granule which 

 has given rise to the blepharoplast of the new flagellum, as 

 described above. These granules have arisen by division of the 

 bud which came from the endosome in the early prophase. The 

 rhizoplast which marks the path of origin of the bud from the 

 endosome may be seen as a persistent structure in one of the 

 nuclei. From each granule can be seen a rhizoplast passing 

 upward to the lower border of the reservoir and ending in the 

 blepharoplast. From the blepharoplast a connecting rhizoplast 

 is seen passing to the basal granule of the smaller branch of the 

 flagellum. From both the blepharoplast and basal granule the 

 new branches of the flagellum may be seen passing upward 

 through the reservoir into the gullet. 



The endosomes of the daughter nuclei become rounded; the 

 chromosomes take on a somewhat spiral arrangement in the 

 peripheral region of each nucleus; condensed nodes appear in 

 each chromosome, giving to them a beaded appearance. Each 

 definite chromosome loses its distinct outlines and gradually 

 becomes a part of the rapidly forming network of chromatin 

 characteristic of the vegetative nucleus. 



During these reorganization stages of the telophase, the 

 chromosomes seem to change their staining capacity. Each 

 appears more diffuse and broader. This appearance may be 

 variously interpreted. In the first place it is conceivable that the 

 change is due to a longitudinal splitting of each chromosome, the 

 halves remaining close together without separation before passing 

 into the resting condition. This is the interpretation which 

 Tschenzoff gives for Euglena viridis. Another explanation is that 

 the metabolic changes in the chromosomes during the reorgani- 

 zation period, especially the relative change from gel to sol, is 

 sufficient to give the diffuse appearance. 



Division of the Gullet and Reservoir. 



As has been mentioned above, the nucleus migrates anteriorly 



during the early prophase and comes into contact with the lower 

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