No. 3.] OESOPHAGEAL GLANDS OF URODELA. 89 



Diese Zellen zeigen in ihrem 1'rotoplasma cinen kornigen Ban, 

 Korner, welchc sich mit verschiedenen Farben, z. B. Eosin, 

 S.-Fuchsin tingieren, mit Osmiumsaure braunen . . . ." No 

 glands resembling the anterior oxyntic glands of Triton are 

 present in the adult, but he found in the young animal, at the 

 junction of oesophagus and stomach, glands which are inter- 

 mediate in nature between the oesophageal and gastric glands. 



In Necturus there are, according to Kingsbury, three kinds 

 of glands present. In the oesophagus are large saccular glands 

 in most respects like those of Proteus, except that Kingsbury 

 was unable, even after repeated trials, to demonstrate the pres- 

 ence of any granules capable of reducing osmic acid. At the 

 junction of oesophagus and stomach are richly branched glands 

 like the anterior oxyntic glands of Triton, and finally there are 

 the ordinary gastric glands. 



There are thus three types of gland occurring in the oesopha- 

 gus of Batrachia, the relationship of which to one another, to 

 the gastric glands, and to the oesophageal glands of higher 

 vertebrates, is obscure. These are the compound pepsin- 

 forming glands of the frog's oesophagus, the saccular glands 

 of the oesophagus of Proteus and Necturus, and the anterior 

 oxyntic glands of Triton and Necturus. It might be claimed 

 for a priori reasons that no possible relationship could exist 

 between the oesophageal glands and the gastric glands, but that 

 position would necessitate a critical examination of the data 

 on which this anatomical division of the foregut in the forms 

 mentioned has been decided. 



The writer found that Amblystoma combined, in a sense, 

 the conditions found in Proteus and Triton, inasmuch as the 

 glands in the larva resemble those of Proteus, the glands of the 

 adult those of Triton. The present memoir is a brief account 

 of the histogenesis of the glands in question. 



Before passing on to a consideration of the histogenetic 

 phenomena it is necessary to describe briefly the structure of 

 the mucous membrane of the foregut in the adult animal. 



The oesophagus is non-glandular, and is lined throughout 

 by a ciliated epithelium, in which many goblet cells may be 

 recognized. The ciliated epithelium is succeeded by the 



