96 BENSLEY. [VoL. II. 



backward, the glands become gradually tubular or saccular, 

 without any appearance of distention, and the ciliated cells dis- 

 appear, so that the rest of the glandular portion, as well as all 

 the posterior non-glandular portion of the stomach, is provided 

 with a mucigenous epithelium. 



Attention should be called at this stage to the remarkable 

 resemblance between the mucigenous border of the gastric 

 epithelium and the cuticula of the cells in the buccal cavity. 

 Both have a characteristic striated appearance, and one is 

 tempted to think that they cannot be very different chem- 

 ically. 



The cells of the tubular glands do not differ in any respect 

 from those of the flask-shaped glands. The mucous cells are 

 less numerous, and a few glands may be entirely without them. 

 The cells of all the glands, even the very last, contain both 

 zymogen granules and prozymogen. 



There are as yet no pyloric glands formed ; the epithelium 

 of the posterior portion of the stomach is perfectly smooth 

 and without glandular outgrowths. 



Even at this stage there is a remarkable resemblance be- 



o 



tween the anterior flask-shaped glands and the oesophageal 

 glands of Proteus and Necturus, and as development proceeds 

 this resemblance becomes more and more striking. 



The mouth and pharynx are lined in the aquatic Amblystoma 

 larva by a stratified non-ciliated epithelium, with cuticular cells 

 and goblet cells. In a transverse series it may be seen that 

 immediately behind the last gill slit this changes to a ciliated 

 epithelium. One may thus consider the first ciliated cell in a 

 longitudinal section as indicating where the oesophagus begins. 

 Measured from this point the foregut in a larva 16 mm. long 

 is about 3 mm. in length. Of this .49 mm. at the anterior 

 end is non-glandular. Behind this we have a portion .45 mm. 

 long extending from the anterior border of the first gland to a 

 point where the foregut begins to expand to form the stomach. 

 This would doubtless, but for the presence of glands, be regarded 

 as a portion of the oesophagus. Beyond this again, the ciliated 

 epithelium extends into the stomach for a distance of .35 mm. 

 The rest of the stomach is lined by a mucigenous epithelium, 



