THE PHYLOGENY OF THE TERMITES. 



derive the termites from the Embiidae as the Embiids from the 

 termites. The wings of the Embiids are specialized in an entirely 

 different direction, the legs are more highly developed, as are also 

 the abdominal appendages, the ocelli have disappeared, etc. 

 With respect to their entire organization the termites may be de- 

 rived without difficulty from the Blattids, which agree with them 

 also in the position of the legs and in having large coxse. In- 

 teresting is the existence of termite larvae with prothoracic dila- 

 tations, which have no particular function and ultimately disap- 

 pear." 



In this paragraph Handlirsch clearly formulates a view which 

 seems to have been gaining ground ever since Hagen, nearly 

 half a century ago ('58, p. 31) insisted on the close affinity of 

 the Termitidae with the Blattidae, or cockroaches. 



At this point the subject has been taken up in an interesting 

 paper by Desneux ('04), who has actually found in a North 

 Australian termite (Mastotermes darwiniensis Froggatt) a form, 

 which, though a true termite, is nevertheless structurally transi- 

 tional between the Blattidae and the other termites. This form, of 

 which only the imago is known, is regarded by Desneux as rep- 

 resenting a new subfamily (Mastotermitinae) and leads him to 

 propose the following arrangement of the Isoptera, or Termitidae : 



Isoptera, or 

 Termitidse 



Termitinse 



I Termitini (Tervies). 

 Rhinotermitini {Rhinotermes -\- Arrhinotermes*). 



Hodotermitini 



Calotermitince 



Calotermitini 

 Calotermes. Hodotermes, -\- Podoterm.es 



-\- Stolotermes. 

 Termopsis. 



Mastotermitinse j Mastotermes. 



The blattoid characters of Mastotermes, as determined by 

 Desneux, are the following : The wings have the typical blattoid 

 neuration, both pairs have a distinct anal area, small and bounded 

 by an arched groove in the fore wing, but in the hind wing 

 highly developed and folded during repose. In the anterior 

 wing the basal corneous portion is considerable. The neuration 



