in Hybridisation 55 



character*. In which of the two significations it appears 

 in each separate case can only be determined by the follow- 

 ing generation. As a parental character it must pass over 

 unchanged to the whole of the offspring; as a hybrid- 

 character, on the other hand, it must observe the same 

 behaviour as in the first generation. 



THE SECOND GENERATION [BRED] FROM THE HYBRIDS. 



Those forms which in the first generation maintain the 

 recessive character do not further vary in the second 

 generation as regards this character ; they remain constant 

 in their offspring. 



It is otherwise with those which possess the dominant 

 character in the first generation [bred from the hybrids]. 

 Of these two-thirds yield offspring which display the 

 dominant and recessive characters in the proportion of 

 3 to 1, and thereby show exactly the same ratio as the 

 hybrid forms, while only one-third remains with the domi- 

 nant character constant. 



The separate experiments yielded the following results :- 



Expt. 1. Among 565 plants which were raised from 

 round seeds of the first generation, 193 yielded round seeds 

 only, and remained therefore constant in this character ; 

 372, however, gave both round and angular seeds, hi the 

 proportion of 3 to 1. The number of the hybrids, therefore, 

 as compared with the constants is 1'93 to 1. 



Expt. 2. Of 519 plants which were raised from seeds 

 whose albumen was of yellow colour in the first generation, 

 166 yielded exclusively yellow, while 353 yielded yellow 



* [This paragraph presents the view of the hybrid-character as 

 something incidental to the hybrid, and not " transmitted " to it a 

 true and fundamental conception here expressed probably for the 

 first time.] 



