GOLGI APPARATUS IN STRIATED MUSCLE. 115 



muscle fibers. This matter has in fact received no very extensive 

 examination, but several workers, and their opinion is concurred 

 in by Duesberg ('10 and 'n), have identified the Q- and J- 

 granules as the mitochondrial constituents of striated muscle. 

 But the staining behavior of these bodies changes rapidly in a 

 manner not characteristic of chondriosomes and the evidence 

 that they are of such a nature seems to depend on what might 

 be termed an argument from logical necessity, viz., these bodies 

 sometimes stain like chondriosomes and they have some appear- 

 ance of being the only granules available which could be so 

 identified. As a matter of fact, a figure of heart muscle given 

 by the Lewises ('24, Plate VI., Fig. b), and Duesberg's ('10) 

 Fig. 24 of the voluntary muscle of the fowl, seem to indicate 

 that the mitochondria are really not the Q- or J-granules at all, 

 but granules with a quite different disposition. The matter 

 evidently is one to be analyzed further, using some basis of 

 identification other than that of staining reactions. 



In conclusion, therefore, my purpose in this short resume has 

 been to bring out our uncertainty concerning some very essential 

 structures in striated muscle fibers ; at the same time to indicate 

 the probable disposition of the Golgi apparatus, and to point 

 out various possibilities which this disposition at once suggests. 

 We have now to reexamine the whole problem in order to establish 

 the homology of the network with the Golgi material of embryonic 

 cells; and to determine the true significance of the Q- and J- 

 granules whether chondriosomes, possible secretory granules or 

 some as yet unrecognized combination of the two. The solution 

 of these problems should throw much interesting light on the 

 functional significance of the Golgi apparatus, and perhaps also 

 of the mitochondria. 



LITERATURE CITED. 

 Cajal, R. y. 



'90 Coloration par la methode de Golgi des terminaisons des trachees et des 



nerfs dans les muscles des insectes. Zeitsch. f. vviss. Mikr., Vol. 7. 

 Duesberg, J. 



'10 Les chondriosomes des cellules embryonnaires du poulet et leur role dans le 

 genese des myofibrilles. Arch. f. Zellforsch., Vol. 4. 



'n Plastosomen, " Apparato reticolare interno," und Chromidialapparat. 

 Anat. Hefte, 2. Abt., Vol. 20. 



