46 SHELLEY R. SAFIR. 



The four classes of males taken together number 329, which 

 approximates to the 345 red females. The results may be 

 accounted for in the following way, using the new system of 

 symbols proposed by Morgan (1912). 



red .......... 9 XCV XCV 



p 



cherry verm.. . cf Xcv 



Fi FEMALE. Fi MALE. 



f XCV f XCV 



{ Xcv \ - 



red 9 red c? 



Gametes of Fi 9 



XcV XCV Xcv XCv 

 Gametes of Fi cf 



XCV - 



F- FEMALES. F 2 MALES. 



XcV XcV , 



red 9 _ cherry cf 



XCV , XCV 



red 9 _ red 



Xcv . ,-. Xcv , - 



red 9 cherry verm, cr 



red 9 verm, rf 



It will be noted that the red-eyed females belong to four classes 

 pure red, reds heterozygous for vermilion, reds heterozygous for 

 cherry, and reds heterozygous for vermilion and cherry. The red 

 and cherry vermilion male classes are each about twice as numer- 

 ous as the cherry and the vermilion classes respectively. This 

 disturbance in the ratio is due to linkage between cherry and 

 vermilion. 



CHERRY VERMILION BY VERMILION. 



When vermilion-eyed females were mated to cherry vermilion- 

 eyed males all the offspring, 483 in number, were vermilion. 

 These when inbred produced vermilion females, vermilion males 

 and cherry vermilion males. 



f verm 9 2<2\ f verm ' ? ......... 374 



verm. 9 by cherry verm, c? = \ JSi > 1 verm, cf ......... 158 



I verm, cr 23i/ ,, 



{ cherry verm, cr ... 158 



The results bear out the view that the new eye color repre- 

 sented the double recessive; vermilion plus some other sex- 

 linked factor, viz., cherry, as the analysis shows: 



verm. 9 . .XCv XCv 



cherry verm, cf Xcv 



