HIPPODAMIA. 



39 



of the progeny have a few small spots. None were fully spotted. Com- 

 bining the progeny of these 12 (81 in all) and weighting small, hazy transi- 

 tional spots as one-half, we find spots 1 present 4, spot 6 present 5 + I, 

 spot 4 present 9 + |, spot 3 present 8 -|- , spot 2 present 11 + f , spot 

 5 present 16 + f . This higher degree of persistence of spots 2 and 5 and 

 the lower degree of 1 and 6 we have also seen in the variation results in 

 fig. 18. Unfortunately I have no cross of typically spotted and spotless. 

 In 149 we have spotless x ; spots 3 and 5. The progeny showed these 

 spots in 6 + iand 6-j-|, respectively, in the 11 offspring. A spotless 

 individual never gave fully-spotted offspring, nor did fully-spotted indi- 

 viduals ever give spotless offspring. Partly-spotted patterns can not be 

 considered a heterozygous condition, because in one experiment where 

 the parents (387) were 2 partly spotted individuals, the 20 offspring 

 were spotless or partly so. The spots which were present most fre- 

 quently in the offspring were present in the mother, but the particular 

 degree of spottedness was not stable, for several were wholly spotless 

 and several were provided with more and better spots than the parents. 

 We may conclude that in spottedness : ( spotlessness simple Mendelian 

 heredity does not prevail, but that crossing gives an intermediate con- 

 dition which, while not acting like a heterozygote, is yet unstable and 

 contributes to both conditions. 



TABLE 13. Percentage of progeny to inherit a characteristic on simple Mendelian 



expectation. 



Spots 1, 2, and 6 are often solely absent in regions where spotlessness 

 prevails, so that their individual heredity becomes of interest. Spot 6 was 

 not lacking in any of the offspring of No. 228 ?, which was so character- 

 ized, but of the similar No. 232 ? the single offspring had the spot reduced. 

 In No. 455, where 6 was transitional, it was normal in all the offspring. 



