HEREDITY 423 



tion is unseen in them simply because black hides it. The white will 

 reappear among the grandchildren. In Fig. 5 we see a mother guinea- 

 pig having a jet black coat. Beside her are four young of the same 

 color as herself. The father too was black. In a word this black race 

 breeds true. A female of this race was mated with the albino male 

 shown in Fig. 6. Albinos have white hair and pink (unpigmented) 

 eyes, the red eye color being due to the blood which shows through; 

 they breed true among themselves, but the result is very different when 

 they are mated with black individuals. Two children of the albino 

 male and the black female are shown in Fig. 7. They are intensely 

 black pigmented, as are all the young produced by this cross. Two of 

 them when grown to maturity and mated with each other, produced a 

 litter of four young, shown in Fig. 8. Three are black pigmented like 

 the parents, but one is an albino similar in all respects to the albino 

 grandsire. Here we notice the reappearance of the albino character 

 after skipping a generation. The albino grandsire really made a hered- 

 itary contribution as regards the character hair color, but it did not 

 show in the children, because black also was present in the children, 

 and black obscured or dominated the white. 



Applying our principle of single germ, dual individual to this case, 

 we see that the facts observed are fully in harmony with it. The 

 original cross brought together the characters B (black) and W (white) 

 into an individual (or zygote as we call it, a joining together) B W, 

 which showed only black. Two such individuals, a male and a female 

 were now mated together. In the formation of germ-cells by these 

 individuals there is a return to the single condition, B separates from 

 W and passes into a different germ-cell. Accordingly, the mother forms 

 eggs, B and W, respectively, and the father forms sperms of a like 

 character. Now a new individual arises from a union of an egg with a 

 sperm. Apparently either sort of sperm may unite with either sort of 

 egg which it chances to meet. So there are formed in the next genera- 

 tion three sorts of zygotes (individuals), viz., B B, B W, and WW, 

 instead of B W alone as in the previous generation. The chances for 

 the occurrence of these three sorts of unions are 1 BB, 2 BW, and 

 1 WW. Any individual containing the character B will be black; 

 accordingly the BWs as well as the BBs will be black and there should 

 be three blacks to one white. These are in fact the observed propor- 

 tions. The white individual should transmit no other character, be- 

 cause it contains only W. Such is indeed the observed fact. Any two 

 white individuals mated together will produce only white offspring. 

 But, if our reasoning is correct, two thirds of the black individuals of 

 this generation (viz., the BWs) should transmit white as well as black, 

 while the remaining one third, BB, should transmit only black. Ex- 

 periment justifies both these conclusions. If we mate the black animals 



