THE VITAL PHENOMENA OF THE CELL 



183 



mences. The nuclein granules, which are distributed all over the 

 linin framework (Fig. 75 A), collect together at certain places and 

 arrange themselves into delicate spiral threads, which are covered 



A B 



Fig. 75. A Resting nucleus of a sperm-mother-cell of Salamandra maculata (after 

 Flemming, PI. 23, Fig. 1 ; from Hatschek). B Nucleus of a sperm-mother-cell of Sala- 

 mandra maculata. Coil stage. The nuclear threads are already commencing to split 

 longitudinally (diagrammatic, after Flemming, PI. 2G, Fig. 1 ; from Hatschek). 



with small indentations and swellings. From these, innumerable 

 most delicate fibrils branch off at right angles ; these fibrils, which 

 consist of strands of the linin framework, only become visible as 

 the nuclein withdraws itself from their surface. Later on the 

 nuclein threads become still more clearly defined, and, as the in- 

 dentations and swellings disappear, develop a perfectly smooth sur- 

 face (Fig. 75 B). Now since they 

 surround the nuclear space on 

 every side, they produce an ap- 

 pearance described by Flemming 

 as the coil figure (spirem, skein). 

 The coil is much more dense in 

 the epithelial cells of Salamandra 

 than in sperm cells, whilst at the 

 same time the threads are much 

 finer and longer (Fig. 76). 



It is as yet undecided, whether 

 at the outset the coil consists of a 

 single long thread or of several 

 such threads. I agree with Rabl 

 (VI. 53) that the latter is more ^ _. 



Fig. 76. Nucleus of an epithelial cell 

 probable, at the commencement of division; from 



A Striking difference is llOW seen a Salamander larva. Fine coil forma- 



, tion. The remains of two nucleoli are 



in the way the various nuclear stl n present. (After Flemming.) 



