462 PROCEEDINGS OF THE ACADEMY OF [1896. 



in the Ophidia, by the presence of calyculi. Such are characteris- 

 tic of the Rhiptoglossa and Pachyglossa. The Nyctisaura possess 

 the same feature. The Diploglossa, Helodermatoidea and Theca- 

 glossa have the organ flounced, the flounces often pocketed or 

 repand on the margin. In the Leptoglossa we have laminae only ; 

 in the Tiid^ mostly transverse, and in the Scincidae mostly longi- 

 tudinal. In various genera terminal papillre are present. The 

 organ may be simple or bifurcate or merely bilobate. I have not 

 met with the case so common in Ophidia, where the sulcus spermati- 

 cus is bifurcate and the organ undivided. 



The structures of the hemipenis have a constant systematic value. 

 As in the Ophidia, the value differs with the character, but it varies 

 from generic to superfamily in rank. 



In the Chamaeleonidfie the greater part of the surface of the hem- 

 ipenis is coarsely calyculate, generally in a transvere direction. 

 There are remarkable papillae at the apex, which differ in the differ- 

 ent forms. In C. 2)ardalis there is a kind of membranous apron 

 proximad of the papillse which presents an apex proximad opposite 

 to the sulcus spermaticus. In C. vulgaris and C. gracilis the papil- 

 lae are erect, laminiform and transverse and serrate on the edges. 

 The principal pair have a few papillse in front of and behind them, 

 and in C. gracilis there is, behind these, on each side, an oval l)ody 

 which is composed of three serrate laminse packed obliquely together. 

 In C. gracilis the proximal lamiuse are low and have a margin of 

 acute tubercles, and each serves as a collar to a much larger papilla. 

 The latter is largely free and tongue-shaped, with the apex proxi- 

 mad, and its flat external surface is covered with three or four rows 

 of conic papillse. 



I have had the opportunity of examining the hemipenis of a 

 relatively small number of species of the Agamidae ; the surface is 

 generally calyculate. I have not found terminal papillre in the 

 genera Uromastix, Agama, Liolepis, Pkysignathus or Calotes. The 

 general construction is, that opposite the sulcus spermaticus is a 

 strong longitudinal welt. Near the apex this welt becomes adher- 

 ent to the side on which the sulcus runs, dividing the organ into two 

 apical portions. The sulcus bifurcates and passes along the base of 

 this partition. In Liolepis there are two welts enclosing a smooth 

 space between them. In Calotes cristatellus there is a lesser welt on 

 each side of the principal one. In all the genera the basal part is 

 smooth, and it is sometimes thrown into longitudinal folds. 



