294 



SUMMARIES OF THE HISTOLOGIC CHARACTERS, ETC. 



polariscopic reactions both incline to N. bowdeni, 

 but N. abundance is not so close as N. giantess. 

 In the iodine reactions with the raw grains the 

 hybrids are as well separated from each other 

 as they are from the parents. In N. giantess the gela- 

 tinized grains behave more like those of N. bou'deni, 

 while the raw grains lean to the other parent; but in 

 the other hybrid there was not found any difference 

 in the parental inclinations of both gelatinized and raw 

 grains. The qualitative reactions with the chemical 

 reagents show curious differences, N. giantess in only two 

 of the six reactions inclining to N. bowdeni and in the 

 other four to the other parent; while the other hybrid 

 inclines all six reactions to N. bowdeni. In the reac- 

 tions of chloral hydrate, potassium sulphocyanate, and 

 sodium salicylate N. abundance is closer than N. giantess 

 to N. bowdeni; and in the potassium-sulphocyanate reac- 

 tion the hybrids are closer to each other than to either 

 parent. In the nitric-acid reaction N. giantess is closer 

 to 2V. sarniensis var. corusca major than is 2V. abundance 

 to N. bowdeni, but the hybrids themselves are very close. 

 In the potassium-iodide reaction 2V. giantess leans to 

 N. sarniensis var. corusca major, while the other hybrid 

 inclines to the other parent; but the hybrids are closer 

 to each other than is either to the parent to which it 

 is the more closely related. The quantitative and quali- 

 tative reactions show most interesting differences and 

 independence. 



It will be seen by an examination of the preceding 

 table how variable and absolutely unpredictable is the 

 shifting of hybrid properties toward one or the other 

 parent. Biparental inheritance in each of the designa- 

 tions is manifest ; but in some instances hybrid and 

 parents are very closely alike, and in others the hybrids 

 are more alike or more different than are the parents, or 

 they differ more from the parents or resemble more 

 closely one or the other parent than do the parents them- 

 selves appear to be the same or different. With the first 

 pair of hybrids, 2V. dainty maid inclines in the histologic 

 properties and qualitative reactions, with the exception 

 of the character and arrangement of the lamellae, in every 

 designation to 2V. elegans: while its mate, 2V. queen of 

 roses, leans in only about two-thirds of the designations 

 to the same parent. With the second pair, 2V. giantess 

 inclines in about one-half of the designations to 2V. bow- 

 deni, while 2V. abundance inclines almost wholly to the 

 same parent. With the last hybrid, 2V. glory of sar- 

 nia, the inclination with the exception of a single desig- 

 nation is to 2V. sarniensis var. corusca major. Excess 

 and deficit of development are rarely noted, and no indi- 

 viduality of the hybrid in any case was recorded. In the 

 quantitative reactions there is obvious independence of 

 the qualitative reactions, inasmuch as they may or may 

 not correspond. In N. dainty maid, while in both histo- 

 logic properties and qualitative reactions the inclination 

 is positively to the pollen parent, in the quantitative 

 reactions there is a tendency to intermediateness, and 

 to the pollen parent. In 2V. queen of roses there is an 

 inclination of about two-thirds of the histologic proper- 

 ties and qualitative reactions to the pollen parent, while 

 in the quantitative reactions there is more of a leaning 

 to the pollen than to the seed parent. In 2V. giantess 

 about one-half of the histologic properties and qualitative 

 reactions lean to the seed parent, in the quantitative 



reactions six of the eight reactions lean to the pollen 

 parent. In 2V. abundance the histologic properties and 

 qualitative reactions incline almost wholly to the seed 

 parent, in the quantitative reactions six of the eight in- 

 cline to the pollen parent. In 2V. glory of sarnia the 

 histologic properties and qualitative reactions incline 

 almost wholly to the seed parent and the quantitative 

 reactions incline equally to each of the two parents. 



NARCISSUS. (TABLE C 6.) 



The first two hybrids, while showing throughout the 

 various designations biparental inheritance, usually bear 

 a closer relationship to 2V. poeticus poetarum than to 

 2V. poeticus ornatus; and on the whole are closer to one 

 another than to either parent. It is strange that while 

 2V 7 . poeticus herrick is in form, hilum, and lamellae closer 

 to 2V. poeticus ornatus than to the other parent, the rela- 

 tionship in size and all other designations is closer to 2V. 

 poeticus poetarum. 2V. poeticus dante is in form closer 

 to 2V. poeticus ornatus, but in all ottier designations 

 closer to the other parent. In form both hybrids are 

 closer to 2V. poeticus ornatus, but 2V. poeticus herrick 

 is the closer of the two. In hilum and lamellae, 2V. poeti- 

 cus herrick shows as close relationship to 2V. poeticus 

 ornatus as does 2V. poeticus dante to 2V. poeticus poe- 

 tarum. In size, 2V. poeticus herrick is closer than 2V. 

 poeticus dante to 2V. poeticus poetarum. In both polari- 

 scopic figure and selenite reactions both hybrids are 

 closer, and in equal degree, to 2V. poeticus poetarum. 

 In the iodine reactions the hybrids do not differ and are 

 therefore equally close to 2V. poeticus poetarum. 

 Throughout the qualitative chemical reagent designa- 

 tions the hybrids are closer to 2V. poeticus poetarum. 

 In the chloral-hydrate and nitric-acid reactions 2V. poeti- 

 cus dante is closer than 2V. poeticus herrick to 2V. poeti- 

 cus poetarum; but in the chromic-acid and pyrogallic- 

 acid reactions the reverse. Only rare records of deficient 

 development were recorded ; in no instance was there 

 excess of development or individuality. In the quanti- 

 tative reactions 2V. poeticus herrick is mid-intermediate 

 or shows a closer relationship to the pollen parent ; while 

 2V. poeticus dante is mid-intermediate in three of the 

 seven reactions and shows a closer relationship in two 

 to the seed parent, and in two to the pollen parent. It 

 is of interest to note that while in the qualitative reac- 

 tions both hybrids are throughout very much closer to 

 the pollen parent than to the seed parent, in the quantita- 

 tive reactions the first leans markedly to the pollen parent 

 and the second to one as much as to the other parent. 



There is seen throughout the designations of the 

 various sets of Narcissi the same swinging of hybrid 

 development to one or the other parent, the independence 

 of each unit-character and unit-character-phase of every 

 other in its direction and degree of development, the 

 absolute impossibility of forecasting the parental rela- 

 tionship of any designation, and the usually close rela- 

 tionship of the hybrid in its properties, as a whole, to 

 one or the other parent, as is evident in preceding sets. 

 Special features of the Narcissi group are attached to 

 the relative potencies of certain of the parents that occur 

 in a number of sets, and to the hybrid 2V. ma.Aa.ine de 

 graaff, which in two sets is the pollen parent. 2V. poeti- 

 cus ornatus is the seed parent in Set 1 and the pollen 

 parent in Sets 2, 3, and 4. As the seed parent, it exhibits 



