158 POPULAR SCIENCE MONTHLY. 



resulting crop of fruit gave no visible indication of impending muta- 

 tion. I as carefully preserved seed from this crop as I had done in 

 the former case, and planted them in my garden in 1901, believing 

 that I should produce Acme plants, notwithstanding my former ex- 

 perience. On the contrary, the result was an exact duplication of 

 my experience with the crop of 1899, every plant and every fruit par- 

 taking fully and uniformly of the duplicated mutation. The plant 

 description, including that of the fruit, which is given in the im- 

 mediately preceding paragraph applies as exactly to the plants of the 

 crop of 1901 as it does to those of the crop of 1899. Fig. 1 represents 

 a plant of this crop in the early stage of its growth, when it was 

 beginning to shed its first flowers. Its deeper shade of green adds to 

 the difference of aspect between the mother and daughter forms. 



The Figures 1 and 2 are copies of photographs which were taken 

 of the plants as they were then growing in my garden.* The plant 

 represented by Fig. 1 bore the new variety, which I have called the 

 Washington. That which is represented by Fig. 2 bore the Acme 

 variety. Although it can not be proved that the particular plant 

 which is represented by Fig. 1 actually came from a seed borne by 

 the plant represented by Fig. 2, I do not hesitate to assert positively 

 that the plant form represented by Fig. 1 is the immediate progeny 

 of the form represented by Fig. 2. I make this statement with all 

 the more confidence because all the work of my garden has been done 

 con amore by my own hands, including the planting of the seed, the 

 plucking of the fruit from which the seed was taken, and the curing 

 and preserving of the seed for the next year's planting. In all this 

 work I practised the same conscientious care that I have done in all 

 my other scientific work in other fields. No tomato seed other than 

 that which I have mentioned was in my possession during all the 

 time my experiments were in progress, and I do not admit the pos- 

 sibility that any other seed was at any time substituted. Even if 

 there had been any such substitution, it would not account for the 

 mutations which I have described, which were phylogenetic in char- 

 acter and not the result of hybridization. The fruit of the mutated 

 plant species was also a new variety and would not, in any ordinary 

 case of germination, have been produced by seed of any other variety 

 previously known. This new variety is as distinct as are any other 

 fine varieties, and it has been true to seed every year since its origina- 

 tion. If my Acme plants, in either of the cases mentioned, had re- 

 ceived adventitious fertilization by pollen from any other flowers than 

 those of their connate crop associates, the cross fertilization would 



* These figures were originally published in an article by Dr. R. France", in 

 Die Umschau, at Frankfurt am Main. In that article it was unfortunately 

 stated that Fig. 1 represents the mother form, while the reverse is the fact. 



