TAXONOMY NYMPHAEA TETRAGONA. 



169 



of anthers as in median stamens. Pollen yellow, finely tuberculate, operculum smooth ; 

 diameter 0.03 to 0.04 mm., average 0.034 mm. Carpels 6 to 8 (4 to 12, as hitherto 

 described) ; styles thin, subcoriaceous, ovate or oblong-elliptic, short and blunt, breadth: 

 length = i : 1.25 to 3, concave; stigmatic area extending on their bases as an upraised 

 cushion, ending obtusely. Stigma little or not at all depressed. Fruit spherical, naked, 

 green, crowned with the small stigma. Seed large, 0.3 to 0.2 cm. long by 0.16 to 

 0.17 cm. in diameter, dark brown, smooth. 



Seeds which germinate in late summer form small plants, and at approach of 

 winter, lose their leaves and retire into a small tuber. These germinate, producing 

 first a series of submerged leaves, then small floating leaves as follows : Submerged 

 leaves broadly obovate, rounded at apex, thin, pale green ; sinus deep, margins nearly 

 straight near the petiole and parallel or diverging or slightly overlapping, curved out- 

 ward below to the broadly rounded lobes. First floating leaf obovate-oblong, rounded 

 at apex ; sinus overlapping, lobes narrow, rounded at angles, suggesting rabbit-ears ; 

 green above with a brown line around margin; under surface greenish-red, densely 

 marked with fine bright red marblings. 



Leaf of mature plant cleft to the petiole, more or less coriaceous, broadly oblong, 

 ovate or elliptic, 5 to 10 cm. long by 3.3 to 7.6 cm. wide, entire, rounded at apex 

 (rarely emarginate, Casp. 1865), dark green above, more or less mottled (at least when 

 young) with reddish-brown ; under surface green to dull red ; primary veins 5 or 6 on 

 each side of leaf, sunk in the tissues; length of principal area: radius of leaf = i : 1.5 

 to 1.9 (average i: 1.3). Sinus depth: length of leaf = i : 1.8 to 2.2 (average 1:2), 

 lobes unequal, margins nearly straight or slightly convex, touching, or diverging and 

 1.9 to 3.8 cm. apart at periphery, angles more or less produced, rounded to sub-acute 

 or rarely acuminate. Astomatic area very small ; stomata larger and not so crowded 

 as in Eu-castalia, scattered toward the point of insertion of the 

 petiole ; cells of upper epidermis large. Petiole smooth, terete, 

 very slender, 0.24 to 0.4 cm. in diameter, 10 to 30 cm. long, 

 with 4 main air-canals, 2 larger and 2 smaller, with 4 still 

 smaller ones anteriorly and posteriorly, and one on each side ; 



2 double bundles are present (i anterior and I posterior) with 



3 or 4 smaller ones on either side ; stellate cells projecting into 

 the 6 larger air-canals, rays very slender, long, densely covered 

 with small crystals, quite brittle. Stipules oblong-ovate or 

 elongate-triangular, acute, not at all keeled. 



Rhizome erect, 7 to 15 cm. long, blackish, covered with long hairs (1.3 cm. long) ; 

 apex covered with leaf-bases and stipules. Entirely without branches or offsets. 



LEAVES FROM SMALL SEEDLING-TUBER (IN CENTIMETERS). 



(AFTER CASPARY, 1865). 



a Overlapping. 



